Spodoptera roseae Schaus 1923

Spodoptera roseae (Schaus)
Figs. 27, 94, 140, 301-303, 337, 367, 399, 431, 463, 543
Trachaea [sic] roseae Schaus 1923: 33. Laphygma roseae; Richards 1941: 239.
Spodoptera roseae; Linsley & Usinger 1966: 160; Hayes 1975: 176; Todd & Poole 1980: 733; Poole 1989: 925.
Diagnosis. — S. roseae is endemic to the Galapagos Islands and cannot be confused with other species found there. The male antenna is serrate
form antennae (Fig. 18). Forewing ground color is brown with a brown patch along the subterminal line. The male genitalia have the ampulla extending to just below the apex of the valve (Fig. 367); coremata in form of a double lobe with dorsal lobe more than half the length of ventral lobe. The distal margin of the ventral plate of the ostium bursa is slightly concave (Fig. 463).
Description. Adult Male. — Head: Basal segment of labial palpus black; median segment dark gray to two-thirds length, apical third mixed rufous and cream (apical third can be light gray); apical segment dark gray. Frons with short scales rufous and cream; long scales rufous; lateral spots at eye margin black; vertex brown and cream. Antenna serrate; scape cream and brown; flagellum cream and brown.
Thorax: Patagium brown with thin white apical margin. Tegula brown (some scales tipped white). Mesothorax brown with some scales tipped white. Underside of thorax cream. Prothoracic femur fuscous; tibia with lateral scale tufts not extending beyond first tarsal segment, fuscous and cream, subapical spot absent; tarsal segments fuscous with cream apical rings. Mesothoracic femur cream and rufous; tibia cream and rufous, inner spur less than twice length of outer spur, internal surface at base thinly devoid of scales on large spur; tarsal segments fuscous with cream apical rings. Metathoracic femur cream and rufous; tibia cream and rufous; tarsal segments fuscous with cream apical rings.
Forewing: Length, 20-22 mm. Ground color dark grayish brown; basal line absent; short longitudinal black dash at base of Cu vein; R vein not a contrasting color; basal scale patch between R and Cu veins absent; basal half circle scale patch below Cu vein absent; antemedial line black; claviform spot absent; basal scale patch absent; gray patch covering anal cell and half of cell CuA2 absent; oblique brown mark from fifth outer marginal spot to vein CuA2 absent; curved white line from antemedial line to postmedial line absent; orbicular spot round, cream, with buff to brown center, outlined in black; reniform spot cream out
lined in black, center light brown to brown; white scales along Cu vein and bases of M3 and CuAl veins absent; postmedial line black, from just below costa to posterior margin, wavy; postmedian band with an indistinct, round, black spot along subterminal line; subterminal line white, indistinct; apex lacking a white patch; black trapezoidal-shaped spots along outer margin; fringe brown. Underside ground color pale gray; outer margin with black spots; reniform spot darker, indistinct.
Hindwing: Ground color pale gray; gray scales along apex and outer margin; fringe white with gray median line to vein CuA2. Underside ground color pale gray; costal cells C and Sc white speckled with gray scales; outer margin with gray triangular spots in middle of cells Sc to CuAl; no spot on underside.
Abdomen: Dorsum pale gray; venter pale gray; disto-lateral black spots on sternites; 8th tergite with spiculate patches present on caudal margin.
Genitalia: Uncus evenly curved; slender, gradually narrowing toward pointed apex; setae absent or few. Scaphium elongate, weakly developed. Tegumen lacking a pair of projecting arms on upper third. Costa straight (Figs. 337, 367). Costal process narrow, elongate; at base of costa on inner edge; perpendicular to costa of valve. Cucullus apex truncate; not produced. Anellifer with lightly sclerotized plate bearing setae present. Ampulla elongate, slightly curved with a decurved apex; extending to just below apex of valve (Figs. 337, 367). Clasper proper absent. Clavus short, round, covered with setae. Sacculus widest at one-third its length, distal half narrow, constant width; apex truncate. Valvula wider than valve; well differentiated from valve, apex free; apex truncate; indentation narrow, deep, ventral margin slightly curved. Coremata elongate, more than 0.5 length of valve; in form of a double lobe. Juxta a narrow rectangular band, median process moderately broad at base and elongate, ventral margin broadly concave. Anellus with minute granules. Vinculum U-shaped with parallel arms and a robust base. Aedoeagus straight; coecum smaller in diameter than shaft; patch of spines absent on apex of membranous sheath surrounding aedoeagus. Vesica curving ventrally; elongate, greater than length of aedoeagus; apicobasal cornutal patch a wide ribbon; length moderate, extending to before middle of vesica; cornuti in form of minute flat granules; lateral cornutal
patch a wide elongate ribbon; a mixture of granular and large spines at apex; dense cornutal patch subventrally on vesica; distal cornutus an ovate plate with large spines at apex.
Adult Female. — Essentially as described for male except:
Head: Antenna filiform.
Forewing: Length, 20-22 mm.
Abdomen: Fine dense scales on 8th segment black.
Genitalia: Ventral plate of ostium bursa with height greater than width; distal margin slightly concave; ventrolateral invaginated pocket of 8th sternite small, more developed ventrally than laterally. Ductus bursae short (length less than twice width); partially sclerotized. Appendix bursae partially sclerotized. Corpus bursae bulbous, length less than twice width; striate convolutions. Signum in apical half of corpus bursae; elongate, length greater than 1.15 mm; forming greater than a 45 degree angle to vertical axis of corpus bursae.
Larva . — Unknown.
Type material. — Trachaea [sic] roseae Schaus. The holotype 3 [USNM] is labeled: Conway Bay; Indefatigable; Galapagos; April 1 1933. Type No. 26507; U.S.N.M. [red label]. Trachea; roseae; type Schs [hand written in black ink]. 3 genitalia on; slide Aug. 1966; E.L.T. 2341. Genitalia Slide; By ELT 3; USNM 46620 [green label]. Type locality: Galapagos: Indefatigable, Conway Bay.
Material Examined. — Specimens were examined from the following country: Ecuador (Galapagos Islands).
Distribution (Fig. 543). — S. roseae is endemic to the Galapagos Islands where it occurs on the following islands: Isabela, Santa Cruz, Santiago, Marchena, and Pinta.