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Image of Pennsylvania Dingy Ground Beetle
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Pennsylvania Dingy Ground Beetle

Harpalus (Pseudoophonus) pensylvanicus (De Geer 1774)

Scientific Names

Preferred Names

Harpalus (Pseudoophonus) pensylvanicus (De Geer 1774)
Species recognized by EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Harpalus pennsylvanicus
Species recognized by United States Species List
Harpalus pensylvanicus
Species recognized by wikipedia NL, Canada Species List, Mexico Species List, North Atlantic Species List, NCBI, and United States Species List
Harpalus pensylvanicus
Recognized by Harvard Museum of Comparative Zoology
Harpalus pensylvanicus (De Geer 1774)
Species recognized by GBIF classification
Harpalus (Pseudoophonus) pensylvanicus (Degeer 1774)
Species recognized by EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Harpalus (Pseudoophonus) pensylvanicus (De Geer 1774)
Species recognized by Catalogue of Life 7 February 2023 and Dynamic Hierarchy Version 2.2 - Test
Harpalus mormonicus (Casey)
Recognized by NMNH type specimens
Harpalus pennsylvanicus
Species recognized by wikidata_hierarchy tar gz and Global Biotic Interactions
Harpalus pensylvanicus
Species recognized by wikidata_hierarchy tar gz, wikipedia EN, Wikidata Common Names, Wikipedia (inferred records), Wikimedia Commons, Barcode of Life Data Systems, Global Biotic Interactions, Global Biotic Interactions, and Global Biotic Interactions
Harpalus pensylvanicus
Recognized by Alaskan Arthropods
Harpalus pensylvanicus (De Geer 1774)
Species recognized by iNaturalist, Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy, and TreatmentBank

Alternative Names

Carabus pensylvanicus De Geer 1774
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Harpalus inmixtus Casey 1924
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Harpalus longior Kirby 1837
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Harpalus mormonicus Casey 1914
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Harpalus pennsylvanicus Say 1823
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Pseudoophonus pensylvanicus (De Geer 1774)
Not accepted according to EOL Dynamic Hierarchy April 2022
Reference taxon from Carabcat database in Species 2000 & ITIS Catalogue of Life
Carabus pensylvanicus Degeer 1774
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Harpalus inmixtus Casey 1924
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Harpalus longior Kirby 1837
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Harpalus mormonicus Casey 1914
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Harpalus pennsylvanicus Say 1823
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Pseudoophonus pensylvanicus (Degeer 1774)
Synonym according to EOL Dynamic Hierarchy 1.1 and EOL Dynamic Hierarchy 1.1
Carabus pensylvanicus De Geer 1774
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Carabus pensylvanicus De Geer 1774
Synonym according to Catalogue of Life 7 February 2023
Harpalus inmixtus Casey 1924
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Harpalus inmixtus Casey 1924
Synonym according to Catalogue of Life 7 February 2023
Harpalus longior Kirby 1837
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Harpalus longior Kirby 1837
Synonym according to Catalogue of Life 7 February 2023
Harpalus mormonicus Casey 1914
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Harpalus mormonicus Casey 1914
Synonym according to Catalogue of Life 7 February 2023
Harpalus pennsylvanicus Say 1823
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Harpalus pennsylvanicus Say 1823 unavailable name
Synonym according to Catalogue of Life 7 February 2023
Pseudoophonus pensylvanicus (De Geer 1774)
Not accepted according to Dynamic Hierarchy Version 2.2 - Test
Pseudoophonus pensylvanicus (De Geer 1774)
Synonym according to Catalogue of Life 7 February 2023

Common Names

English

Pennsylvania Dingy Ground Beetle preferred
Recognized by Wikidata Common Names

Curated hierarchies for Harpalus (Pseudoophonus) pensylvanicus (De Geer 1774)

EOL Dynamic Hierarchy April 2022

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Harpalus
Harpalus (Pseudoophonus) pensylvanicus (De Geer 1774) (this page)
Harpalus (Cryptophonus) idiotus Bates 1889
Harpalus (Cryptophonus) janinae Jeanne 1984
Harpalus (Cryptophonus) schaumii Wollaston 1864
Harpalus (Harpalus) araraticus Mlynar 1979
Harpalus (Harpalus) arcuatus Tschitscherine 1898
Harpalus (Harpalus) asemus Basilewsky 1946
Harpalus (Harpalus) aterrimus Casey 1914
Harpalus (Harpalus) atratus Latreille 1804
Harpalus (Harpalus) atrichatus Hatch 1949
Harpalus (Harpalus) badakschanus Jedlicka 1956
Harpalus (Harpalus) balli Noonan 1991
Harpalus (Harpalus) bellieri Reiche 1861
Harpalus (Harpalus) brachypterus Tschitscherine 1898
Harpalus (Harpalus) brevicornis Germar 1824
Harpalus (Harpalus) cautus Dejean 1829
Harpalus (Harpalus) chalcopterus Jeannel 1948
Harpalus (Harpalus) chobautianus Lutshnik 1922
Harpalus (Harpalus) contemptus Dejean 1829
Harpalus (Harpalus) crates Bates 1883
Harpalus (Harpalus) demelti Korge 1962
Harpalus (Harpalus) dissitus Antoine 1931
Harpalus (Harpalus) dubius Boheman 1848
Harpalus (Harpalus) ebeninus Heyden 1870
Harpalus (Harpalus) egorovi Lafer 1989
Harpalus (Harpalus) famelicus Tschitscherine 1898
Harpalus (Harpalus) fimetarius Dejean 1829
Harpalus (Harpalus) frater Chaudoir 1876
Harpalus (Harpalus) fuscipennis Wiedemann 1825
Harpalus (Harpalus) glasunovi Kataev 1987
Harpalus (Harpalus) herbivagus Say 1823
Harpalus (Harpalus) heyrovskyi Jedlicka 1928
Harpalus (Harpalus) hirtipes (Panzer 1796)
Harpalus (Harpalus) imerinae Jeannel 1948
Harpalus (Harpalus) impressicollis Jeannel 1948
Harpalus (Harpalus) indicola Bates 1878
Harpalus (Harpalus) italus Schaum 1860
Harpalus (Harpalus) kabakovi Kataev 1987
Harpalus (Harpalus) karakorum Jedlicka 1958
Harpalus (Harpalus) kazanensis Jedlicka 1958
Harpalus (Harpalus) kiritshenkoi Kataev 1990
Harpalus (Harpalus) kunarensis Kataev 1993
Harpalus (Harpalus) laetus Reiche 1843
Harpalus (Harpalus) lateralis Dejean 1829
Harpalus (Harpalus) lethierryi Reiche 1860
Harpalus (Harpalus) lewisii Le Conte 1865
Harpalus (Harpalus) lopezi J. Serrano & Lencina 2009
Harpalus (Harpalus) lugubris Boheman 1848
Harpalus (Harpalus) major (Motschulsky 1850)
Harpalus (Harpalus) manas Kataev 1990
Harpalus (Harpalus) martini Van Dyke 1926
Harpalus (Harpalus) mauritanicus Gaubil 1844
Harpalus (Harpalus) metallinus Ménétriés 1836
Harpalus (Harpalus) metarsius Andrewes 1930
Harpalus (Harpalus) meteorus Basilewsky 1946
Harpalus (Harpalus) microthorax (Motschulsky 1849)
Harpalus (Harpalus) mlynari Kataev 1990
Harpalus (Harpalus) modestus Dejean 1829
Harpalus (Harpalus) morvani Kataev 2001
Harpalus (Harpalus) nanniscus Péringuey 1896
Harpalus (Harpalus) natalensis Boheman 1848
Harpalus (Harpalus) natalicus Péringuey 1896
Harpalus (Harpalus) nyassicus Basilewsky 1946
Harpalus (Harpalus) oodioides Dejean 1829
Harpalus (Harpalus) pachys Jeannel 1948
Harpalus (Harpalus) pulvinatus Ménétriés 1848
Harpalus (Harpalus) raphaili Kataev 1997
Harpalus (Harpalus) reversus Casey 1924
Harpalus (Harpalus) rufiscapus Gebler 1833
Harpalus (Harpalus) salinus Dejean 1829
Harpalus (Harpalus) semipunctatus Dejean 1829
Harpalus (Harpalus) smaragdinus (Duftschmid 1812)
Harpalus (Harpalus) spadiceus Dejean 1829
Harpalus (Harpalus) stevenii Dejean 1829
Harpalus (Harpalus) taciturnus Dejean 1829
Harpalus (Harpalus) tangutorum Kataev 1993
Harpalus (Harpalus) tenuestriatus Jeannel 1948
Harpalus (Harpalus) tithonus Reitter 1900
Harpalus (Harpalus) tjanschanicus Semenov 1889
Harpalus (Harpalus) trichophorus Tschitscherine 1897
Harpalus (Harpalus) zhdankoi Kataev 1990
Harpalus (Loboharpalus) platynotus Bates 1873
Harpalus (Pseudoophonus) actiosus Casey 1914
Harpalus (Pseudoophonus) adenticulatus Huang 1992
Harpalus (Pseudoophonus) capito A. Morawitz 1862
Harpalus (Pseudoophonus) coreanus (Tschitscherine 1895)
Harpalus (Pseudoophonus) erythropus Dejean 1829
Harpalus (Pseudoophonus) hatchi Ball & Anderson 1962
Harpalus (Pseudoophonus) katiae F. Battoni 1985
Harpalus (Pseudoophonus) muciulus Huang 1992
Harpalus (Pseudoophonus) paratus Casey 1924
Harpalus (Pseudoophonus) pseudophonoides Schauberger 1930
Harpalus (Pseudoophonus) rufipes (De Geer 1774)
Harpalus (Pseudoophonus) scabripectus Huang
Harpalus (Pseudoophonus) sericatus Tschitscherine 1906
Harpalus (Pseudoophonus) tridens A. Morawitz 1862
Harpalus (Pseudoophonus) ussuriensis Chaudoir 1863
Harpalus ignobilis Boheman 1858
Harpalus phoeopterus Péringuey 1896
Harpalus praedatorius Gistel 1857
Harpalus serbicus Matits 1910
348 additional siblings truncated for brevity.
See the resource file for a full list.

Catalogue of Life 7 February 2023

Biota
Animalia
Arthropoda
Insecta
Coleoptera
Caraboidea
Carabidae Latreille 1802
Harpalinae Bonelli 1810
Harpalini Bonelli 1810
Harpalina Bonelli 1810
Harpalus Latreille 1802
Harpalus subgen. Pseudoophonus Motschulsky 1844
Harpalus (Pseudoophonus) pensylvanicus (De Geer 1774) (this page)
Harpalus (Pseudoophonus) actiosus Casey 1914
Harpalus (Pseudoophonus) aenigma (Tschitscherine 1897)
Harpalus (Pseudoophonus) alienus Bates 1878
Harpalus (Pseudoophonus) aogashimensis (Habu 1957)
Harpalus (Pseudoophonus) azumai Habu 1968
Harpalus (Pseudoophonus) babai Habu 1973
Harpalus (Pseudoophonus) calceatus (Duftschmid 1812)
Harpalus (Pseudoophonus) caliginosus (Fabricius 1775)
Harpalus (Pseudoophonus) capito A. Morawitz 1862
Harpalus (Pseudoophonus) compar Le Conte 1847
Harpalus (Pseudoophonus) coreanus (Tschitscherine 1895)
Harpalus (Pseudoophonus) davidi (Tschitscherine 1897)
Harpalus (Pseudoophonus) disimuciulus Huang
Harpalus (Pseudoophonus) eous Tschitscherine 1901
Harpalus (Pseudoophonus) erythropus Dejean 1829
Harpalus (Pseudoophonus) faunus Say 1823
Harpalus (Pseudoophonus) fokienensis Schauberger 1930
Harpalus (Pseudoophonus) griseus (Panzer 1796)
Harpalus (Pseudoophonus) hatchi Ball & Anderson 1962
Harpalus (Pseudoophonus) hauserianus Schauberger 1929
Harpalus (Pseudoophonus) indicus Bates 1891
Harpalus (Pseudoophonus) jureceki (Jedlicka 1928)
Harpalus (Pseudoophonus) katiae F. Battoni 1985
Harpalus (Pseudoophonus) liobasis Chaudoir 1868
Harpalus (Pseudoophonus) meghalayensis Kataev 2001
Harpalus (Pseudoophonus) meridianus Andrewes 1923
Harpalus (Pseudoophonus) paratus Casey 1924
Harpalus (Pseudoophonus) pastor Motschulsky 1844
Harpalus (Pseudoophonus) poncei Will 2002
Harpalus (Pseudoophonus) protractus Casey 1914
Harpalus (Pseudoophonus) pseudohauserianus Kataev 2001
Harpalus (Pseudoophonus) pseudophonoides Schauberger 1930
Harpalus (Pseudoophonus) roninus Bates 1873
Harpalus (Pseudoophonus) rufipes (De Geer 1774)
Harpalus (Pseudoophonus) sericatus Tschitscherine 1906
Harpalus (Pseudoophonus) simplicidens Schauberger 1929
Harpalus (Pseudoophonus) singularis Tschitscherine 1906
Harpalus (Pseudoophonus) sinicus Hope 1845
Harpalus (Pseudoophonus) suensoni Kataev 1997
Harpalus (Pseudoophonus) texanus Casey 1914
Harpalus (Pseudoophonus) tridens A. Morawitz 1862
Harpalus (Pseudoophonus) ussuriensis Chaudoir 1863
Harpalus (Pseudoophonus) vagans Le Conte 1865
Harpalus (Pseudoophonus) yinchuanensis Huang 1993

Dynamic Hierarchy Version 2.2 - Test

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Harpalus
Harpalus (Pseudoophonus) pensylvanicus (De Geer 1774) (this page)
Harpalus (Cryptophonus) idiotus Bates 1889
Harpalus (Cryptophonus) janinae Jeanne 1984
Harpalus (Cryptophonus) schaumii Wollaston 1864
Harpalus (Harpalus) araraticus Mlynar 1979
Harpalus (Harpalus) arcuatus Tschitscherine 1898
Harpalus (Harpalus) asemus Basilewsky 1946
Harpalus (Harpalus) aterrimus Casey 1914
Harpalus (Harpalus) atratus Latreille 1804
Harpalus (Harpalus) atrichatus Hatch 1949
Harpalus (Harpalus) badakschanus Jedlicka 1956
Harpalus (Harpalus) balli Noonan 1991
Harpalus (Harpalus) bellieri Reiche 1861
Harpalus (Harpalus) brachypterus Tschitscherine 1898
Harpalus (Harpalus) brevicornis Germar 1824
Harpalus (Harpalus) cautus Dejean 1829
Harpalus (Harpalus) chalcopterus Jeannel 1948
Harpalus (Harpalus) chobautianus Lutshnik 1922
Harpalus (Harpalus) contemptus Dejean 1829
Harpalus (Harpalus) crates Bates 1883
Harpalus (Harpalus) demelti Korge 1962
Harpalus (Harpalus) dissitus Antoine 1931
Harpalus (Harpalus) dubius Boheman 1848
Harpalus (Harpalus) ebeninus Heyden 1870
Harpalus (Harpalus) egorovi Lafer 1989
Harpalus (Harpalus) famelicus Tschitscherine 1898
Harpalus (Harpalus) fimetarius Dejean 1829
Harpalus (Harpalus) frater Chaudoir 1876
Harpalus (Harpalus) fuscipennis Wiedemann 1825
Harpalus (Harpalus) glasunovi Kataev 1987
Harpalus (Harpalus) herbivagus Say 1823
Harpalus (Harpalus) heyrovskyi Jedlicka 1928
Harpalus (Harpalus) hirtipes (Panzer 1796)
Harpalus (Harpalus) imerinae Jeannel 1948
Harpalus (Harpalus) impressicollis Jeannel 1948
Harpalus (Harpalus) indicola Bates 1878
Harpalus (Harpalus) italus Schaum 1860
Harpalus (Harpalus) kabakovi Kataev 1987
Harpalus (Harpalus) karakorum Jedlicka 1958
Harpalus (Harpalus) kazanensis Jedlicka 1958
Harpalus (Harpalus) kiritshenkoi Kataev 1990
Harpalus (Harpalus) kunarensis Kataev 1993
Harpalus (Harpalus) laetus Reiche 1843
Harpalus (Harpalus) lateralis Dejean 1829
Harpalus (Harpalus) lethierryi Reiche 1860
Harpalus (Harpalus) lewisii Le Conte 1865
Harpalus (Harpalus) lopezi J. Serrano & Lencina 2009
Harpalus (Harpalus) lugubris Boheman 1848
Harpalus (Harpalus) major (Motschulsky 1850)
Harpalus (Harpalus) manas Kataev 1990
Harpalus (Harpalus) martini Van Dyke 1926
Harpalus (Harpalus) mauritanicus Gaubil 1844
Harpalus (Harpalus) metallinus Ménétriés 1836
Harpalus (Harpalus) metarsius Andrewes 1930
Harpalus (Harpalus) meteorus Basilewsky 1946
Harpalus (Harpalus) microthorax (Motschulsky 1849)
Harpalus (Harpalus) mlynari Kataev 1990
Harpalus (Harpalus) modestus Dejean 1829
Harpalus (Harpalus) morvani Kataev 2001
Harpalus (Harpalus) nanniscus Péringuey 1896
Harpalus (Harpalus) natalensis Boheman 1848
Harpalus (Harpalus) natalicus Péringuey 1896
Harpalus (Harpalus) nyassicus Basilewsky 1946
Harpalus (Harpalus) oodioides Dejean 1829
Harpalus (Harpalus) pachys Jeannel 1948
Harpalus (Harpalus) pulvinatus Ménétriés 1848
Harpalus (Harpalus) raphaili Kataev 1997
Harpalus (Harpalus) reversus Casey 1924
Harpalus (Harpalus) rufiscapus Gebler 1833
Harpalus (Harpalus) salinus Dejean 1829
Harpalus (Harpalus) semipunctatus Dejean 1829
Harpalus (Harpalus) smaragdinus (Duftschmid 1812)
Harpalus (Harpalus) spadiceus Dejean 1829
Harpalus (Harpalus) stevenii Dejean 1829
Harpalus (Harpalus) taciturnus Dejean 1829
Harpalus (Harpalus) tangutorum Kataev 1993
Harpalus (Harpalus) tenuestriatus Jeannel 1948
Harpalus (Harpalus) tithonus Reitter 1900
Harpalus (Harpalus) tjanschanicus Semenov 1889
Harpalus (Harpalus) trichophorus Tschitscherine 1897
Harpalus (Harpalus) zhdankoi Kataev 1990
Harpalus (Loboharpalus) platynotus Bates 1873
Harpalus (Pseudoophonus) actiosus Casey 1914
Harpalus (Pseudoophonus) adenticulatus Huang 1992
Harpalus (Pseudoophonus) capito A. Morawitz 1862
Harpalus (Pseudoophonus) coreanus (Tschitscherine 1895)
Harpalus (Pseudoophonus) erythropus Dejean 1829
Harpalus (Pseudoophonus) hatchi Ball & Anderson 1962
Harpalus (Pseudoophonus) katiae F. Battoni 1985
Harpalus (Pseudoophonus) muciulus Huang 1992
Harpalus (Pseudoophonus) paratus Casey 1924
Harpalus (Pseudoophonus) pseudophonoides Schauberger 1930
Harpalus (Pseudoophonus) rufipes (De Geer 1774)
Harpalus (Pseudoophonus) scabripectus Huang
Harpalus (Pseudoophonus) sericatus Tschitscherine 1906
Harpalus (Pseudoophonus) tridens A. Morawitz 1862
Harpalus (Pseudoophonus) ussuriensis Chaudoir 1863
Harpalus ignobilis Boheman 1858
Harpalus phoeopterus Péringuey 1896
Harpalus praedatorius Gistel 1857
Harpalus serbicus Matits 1910
347 additional siblings truncated for brevity.
See the resource file for a full list.

EOL Dynamic Hierarchy 1.1

Life
Cellular Organisms
Eukaryota
Opisthokonta
Metazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Pancrustacea
Hexapoda
Insecta
Pterygota
Neoptera
Endopterygota
Coleoptera
Adephaga
Carabidae
Harpalus
Harpalus (Pseudoophonus) pensylvanicus (Degeer 1774) (this page)
Harpalus
Harpalus
Harpalus (Cryptophonus) idiotus Bates 1889
Harpalus (Cryptophonus) schaumii Wollaston 1864
Harpalus (Harpalus) araraticus Mlynar 1979
Harpalus (Harpalus) arcuatus Tschitscherine 1898
Harpalus (Harpalus) asemus Basilewsky 1946
Harpalus (Harpalus) aterrimus Casey 1914
Harpalus (Harpalus) atrichatus Hatch 1949
Harpalus (Harpalus) badakschanus Jedlicka 1956
Harpalus (Harpalus) balli Noonan 1991
Harpalus (Harpalus) bellieri Reiche 1861
Harpalus (Harpalus) brachypterus Tschitscherine 1898
Harpalus (Harpalus) brevicornis Germar 1824
Harpalus (Harpalus) cautus Dejean 1829
Harpalus (Harpalus) chalcopterus Jeannel 1948
Harpalus (Harpalus) chobautianus Lutshnik 1922
Harpalus (Harpalus) contemptus Dejean 1829
Harpalus (Harpalus) crates Bates 1883
Harpalus (Harpalus) demelti Korge 1962
Harpalus (Harpalus) dissitus Antoine 1931
Harpalus (Harpalus) dubius Boheman 1848
Harpalus (Harpalus) ebeninus Heyden 1870
Harpalus (Harpalus) egorovi Lafer 1989
Harpalus (Harpalus) famelicus Tschitscherine 1898
Harpalus (Harpalus) fimetarius Dejean 1829
Harpalus (Harpalus) frater Chaudoir 1876
Harpalus (Harpalus) fuscipennis Wiedemann 1825
Harpalus (Harpalus) herbivagus Say 1823
Harpalus (Harpalus) heyrovskyi Jedlicka 1928
Harpalus (Harpalus) hirtipes (Panzer 1796)
Harpalus (Harpalus) imerinae Jeannel 1948
Harpalus (Harpalus) impressicollis Jeannel 1948
Harpalus (Harpalus) indicola Bates 1878
Harpalus (Harpalus) italus Schaum 1860
Harpalus (Harpalus) kabakovi Kataev 1987
Harpalus (Harpalus) karakorum Jedlicka 1958
Harpalus (Harpalus) kazanensis Jedlicka 1958
Harpalus (Harpalus) kunarensis Kataev 1993
Harpalus (Harpalus) laetus Reiche 1843
Harpalus (Harpalus) lateralis Dejean 1829
Harpalus (Harpalus) lethierryi Reiche 1860
Harpalus (Harpalus) lewisii Le Conte 1865
Harpalus (Harpalus) lopezi J. Serrano & Lencina 2009
Harpalus (Harpalus) lugubris Boheman 1848
Harpalus (Harpalus) major (Motschulsky 1850)
Harpalus (Harpalus) manas Kataev 1990
Harpalus (Harpalus) martini Van Dyke 1926
Harpalus (Harpalus) mauritanicus Gaubil 1844
Harpalus (Harpalus) metallinus Ménétries 1836
Harpalus (Harpalus) metarsius Andrewes 1930
Harpalus (Harpalus) microthorax (Motschulsky 1849)
Harpalus (Harpalus) mlynari Kataev 1990
Harpalus (Harpalus) modestus Dejean 1829
Harpalus (Harpalus) morvani Kataev 2001
Harpalus (Harpalus) nanniscus Péringuey 1896
Harpalus (Harpalus) natalensis Boheman 1848
Harpalus (Harpalus) natalicus Péringuey 1896
Harpalus (Harpalus) nyassicus Basilewsky 1946
Harpalus (Harpalus) oodioides Dejean 1829
Harpalus (Harpalus) pachys Jeannel 1948
Harpalus (Harpalus) pulvinatus Ménetriés 1848
Harpalus (Harpalus) raphaili Kataev 1997
Harpalus (Harpalus) reversus Casey 1924
Harpalus (Harpalus) rufiscapus Gebler 1833
Harpalus (Harpalus) salinus Dejean 1829
Harpalus (Harpalus) semipunctatus Dejean 1829
Harpalus (Harpalus) smaragdinus (Duftschmid 1812)
Harpalus (Harpalus) spadiceus Dejean 1829
Harpalus (Harpalus) stevenii Dejean 1829
Harpalus (Harpalus) taciturnus Dejean 1829
Harpalus (Harpalus) tangutorum Kataev 1993
Harpalus (Harpalus) tenuestriatus Jeannel 1948
Harpalus (Harpalus) tithonus Reitter 1900
Harpalus (Harpalus) tjanschanicus Semenov 1889
Harpalus (Harpalus) trichophorus Tschitscherine 1897
Harpalus (Harpalus) zhdankoi Kataev 1990
Harpalus (Loboharpalus) platynotus Bates 1873
Harpalus (Pseudoophonus) actiosus Casey 1914
Harpalus (Pseudoophonus) adenticulatus Huang 1992
Harpalus (Pseudoophonus) capito A. Morawitz 1862
Harpalus (Pseudoophonus) coreanus (Tschitscherine 1895)
Harpalus (Pseudoophonus) erythropus Dejean 1829
Harpalus (Pseudoophonus) hatchi Ball & Anderson 1962
Harpalus (Pseudoophonus) katiae F. Battoni 1985
Harpalus (Pseudoophonus) muciulus Huang 1992
Harpalus (Pseudoophonus) paratus Casey 1924
Harpalus (Pseudoophonus) pseudophonoides Schauberger 1930
Harpalus (Pseudoophonus) rufipes (Degeer 1774)
Harpalus (Pseudoophonus) scabripectus Huang
Harpalus (Pseudoophonus) sericatus Tschitscherine 1906
Harpalus (Pseudoophonus) tridens A. Morawitz 1862
Harpalus (Pseudoophonus) ussuriensis Chaudoir 1863
Harpalus burmeisteri Giebel 1856
Harpalus ewaldi Giebel 1856
Harpalus ignobilis Boheman 1858
Harpalus phoeopterus Péringuey 1896
Harpalus praedatorius Gistel 1857
Harpalus serbicus Matits 1910
Harpalus whitfieldii Scudder 1900
382 additional siblings truncated for brevity.
See the resource file for a full list.

GBIF classification

Animalia
Arthropoda
Insecta
Coleoptera
Carabidae
Harpalus Latreille 1802
Harpalus pensylvanicus (De Geer 1774) (this page)
Harpalus pennsylvanicus mormonicus Casey
Harpalus pennsylvanicus pennsylvanicus
Harpalus actiosus Casey 1914
Harpalus aenigma (Tschitscherine 1897)
Harpalus agakhaniantzi (Mikhailov 1972)
Harpalus akinini Tschitscherine 1895
Harpalus alienus Bates 1878
Harpalus amputatus Say 1830
Harpalus animosus Casey 1924
Harpalus anisodactyliformis Solsky 1874
Harpalus antonowi (Tschitscherine 1898)
Harpalus anxius (Duftschmid 1812)
Harpalus arnoldii Kataev 1988
Harpalus asemus Basilewsky 1946
Harpalus atratus Latreille 1804
Harpalus atrichatus Hatch 1949
Harpalus attenuatus Stephens 1828
Harpalus australasiae Dejean 1829
Harpalus autumnalis (Duftschmid 1812)
Harpalus calceatus (Duftschmid 1812)
Harpalus caliginosus (Fabricius 1775)
Harpalus capito A. Morawitz 1862
Harpalus caspius (Steven 1806)
Harpalus cautus Dejean 1829
Harpalus circumpunctatus Chaudoir 1846
Harpalus cohni Ball 1972
Harpalus cordifer Notman 1919
Harpalus coreanus (Tschitscherine 1895)
Harpalus corpulentus (Casey 1914)
Harpalus cyanopterus (Tschitscherine 1897)
Harpalus desertus Le Conte 1859
Harpalus dimidiatus (P. Rossi 1790)
Harpalus distinguendus (Duftschmid 1812)
Harpalus ellipsis Le Conte 1847
Harpalus erraticus Say 1823
Harpalus erythropus Dejean 1829
Harpalus faunus Say 1823
Harpalus flavescens (Piller & Mitterpacher 1783)
Harpalus fokienensis Schauberger 1930
Harpalus fraternus Le Conte 1852
Harpalus froelichii Sturm 1818
Harpalus fulgens Csiki 1932
Harpalus fulvicornis Thunberg 1806
Harpalus fulvilabris Mannerheim 1853
Harpalus fuscicornis Ménétriés 1832
Harpalus gisellae Csiki 1932
Harpalus gravis Le Conte 1858
Harpalus griseus (Panzer 1796)
Harpalus hatchi Ball & Anderson 1962
Harpalus herbivagus Say 1823
Harpalus honestus (Duftschmid 1812)
Harpalus inconcinnus Chaudoir 1876
Harpalus indianus Csiki 1932
Harpalus indicus Bates 1891
Harpalus indigens Casey 1924
Harpalus innocuus Le Conte 1863
Harpalus laevipes Zetterstedt 1828
Harpalus laticeps Le Conte 1850
Harpalus lethierryi Reiche 1860
Harpalus liobasis Chaudoir 1868
Harpalus luteicornis (Duftschmid 1812)
Harpalus megacephalus Le Conte 1847
Harpalus melancholicus Dejean 1829
Harpalus mitridati Pliginski 1915
Harpalus modestus Dejean 1829
Harpalus morvani Kataev 2001
Harpalus neglectus Audinet-Serville 1821
Harpalus nigritarsis C. R. Sahlberg 1827
Harpalus obliquus G. Horn 1880
Harpalus obnixus Casey 1924
Harpalus ochropus Kirby 1837
Harpalus opacipennis (Haldeman 1843)
Harpalus paratus Casey 1924
Harpalus picipennis (Duftschmid 1812)
Harpalus plenalis Casey 1914
Harpalus protractus Casey 1914
Harpalus pumilus Sturm 1818
Harpalus rubripes (Duftschmid 1812)
Harpalus rufipalpis Sturm 1818
Harpalus schaumii Wollaston 1864
Harpalus semenowi Tschitscherine 1901
Harpalus setiporus (Reitter 1894)
Harpalus signaticornis (Duftschmid 1812)
Harpalus sinicus Hope 1845
Harpalus sinuatus Tschitscherine 1893
Harpalus smaragdinus (Duftschmid 1812)
Harpalus solitaris Dejean 1829
Harpalus somereni Basilewsky 1946
Harpalus somnulentus Dejean 1829
Harpalus spadiceus Dejean 1829
Harpalus stevenii Dejean 1829
Harpalus subcylindricus Dejean 1829
Harpalus suensoni Kataev 1997
Harpalus szalliesi Kataev & Wrase 1995
Harpalus tadorcus Ball 1972
Harpalus tardus (Panzer 1796)
Harpalus tenebrosus Dejean 1829
Harpalus texanus Casey 1914
Harpalus venator Boheman 1848
Harpalus ventralis Le Conte 1847
Harpalus wohlberedti van Emden 1932
Harpalus xanthopus Gemminger & Harold 1868
360 additional siblings truncated for brevity.
See the resource file for a full list.

Integrated Taxonomic Information System (ITIS) Taxonomic Hierarchy

Animalia
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Pterygota
Neoptera
Holometabola
Coleoptera Linnaeus 1758
Adephaga Schellenberg 1806
Carabidae Latreille 1802
Harpalinae Bonelli 1810
Harpalini Bonelli 1810
Harpalus Latreille 1802
Harpalus subgen. Pseudoophonus Motschulsky 1844
Harpalus pensylvanicus (De Geer 1774) (this page)
Harpalus actiosus Casey 1914
Harpalus compar Le Conte 1847
Harpalus erythropus Dejean 1829
Harpalus faunus Say 1823
Harpalus hatchi Ball & Anderson 1962
Harpalus liobasis Chaudoir 1868
Harpalus paratus Casey 1924
Harpalus poncei Will 2002
Harpalus protractus Casey 1914
Harpalus rufipes (De Geer 1774)
Harpalus texanus Casey 1914
Harpalus vagans Le Conte 1865

NCBI

root
cellular organisms
Eukaryota
Opisthokonta
Metazoa
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Panarthropoda
Arthropoda
Mandibulata
Pancrustacea
Hexapoda
Insecta
Dicondylia
Pterygota
Neoptera
Holometabola
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Harpalini
Harpalus
Harpalus pensylvanicus (this page)
Harpalus aeneus
Harpalus aesculanus
Harpalus affinis
Harpalus amputatus
Harpalus anxius
Harpalus atratus
Harpalus attenuatus
Harpalus aureus
Harpalus autumnalis
Harpalus bellieri
Harpalus calceatus
Harpalus caliginosus
Harpalus capito
Harpalus compar
Harpalus contemptus
Harpalus decipiens
Harpalus desertus
Harpalus dimidiatus
Harpalus discrepans
Harpalus dispar
Harpalus distinguendus
Harpalus ebeninus
Harpalus eous
Harpalus faunus
Harpalus fraternus
Harpalus froelichii
Harpalus fulvilabris
Harpalus fuscipalpis
Harpalus griseus
Harpalus herbivagus
Harpalus hirtipes
Harpalus honestus
Harpalus indianus
Harpalus indigens
Harpalus jureceki
Harpalus katiae
Harpalus laevipes
Harpalus laticeps
Harpalus latus
Harpalus luteicornis
Harpalus microthorax
Harpalus modestus
Harpalus neglectus
Harpalus nigritarsis
Harpalus niigatanus
Harpalus opacipennis
Harpalus picipennis
Harpalus pigmaeus
Harpalus pumilus
Harpalus punctatostriatus
Harpalus puncticeps
Harpalus reversus
Harpalus rubripes
Harpalus rufibarbis
Harpalus rufipalpis
Harpalus rufipes
Harpalus serripes
Harpalus servus
Harpalus sinicus
Harpalus smaragdinus
Harpalus solitaris
Harpalus somnulentus
Harpalus subcylindricus
Harpalus tardus
Harpalus tsushimanus
Harpalus ventralis
Harpalus vicarius
Harpalus wagneri
Harpalus xanthopus
unclassified Harpalus

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Harpalus
Harpalus pensylvanicus (this page)
Harpalus actiosus
Harpalus acupalpoides
Harpalus adenticulatus
Harpalus aeneipennis
Harpalus aenigma
Harpalus aequicollis
Harpalus aesculanus
Harpalus agakhaniantzi
Harpalus agilis
Harpalus akinini
Harpalus alajensis
Harpalus albanicus
Harpalus alexandrae
Harpalus alexeevi
Harpalus alienus
Harpalus alpivagus
Harpalus amarellus
Harpalus amariformis
Harpalus amplicollis
Harpalus amputatus
Harpalus anatolicus
Harpalus angulatus
Harpalus angustipennis
Harpalus angustitarsis
Harpalus animosus
Harpalus anisodactyliformis
Harpalus anxioides
Harpalus apache
Harpalus araraticus
Harpalus arcuatus
Harpalus arnoldii
Harpalus ascetes
Harpalus asemus
Harpalus asphaltinus
Harpalus aterrimus
Harpalus atratus
Harpalus atrichatus
Harpalus atripes
Harpalus attenuatus
Harpalus australasiae
Harpalus autumnalis
Harpalus azumai
Harpalus babai
Harpalus badakschanus
Harpalus baleensis
Harpalus balli
Harpalus basanicus
Harpalus basilewskyi
Harpalus basuto
Harpalus bellieri
Harpalus beneshi
Harpalus bitinctus
Harpalus brachypterus
Harpalus brevicornis
Harpalus brevis
Harpalus breviusculus
Harpalus brunnipes
Harpalus bucharicus
Harpalus bungii
Harpalus caeruleatus
Harpalus caiphus
Harpalus calathoides
Harpalus calceatus
Harpalus capicola
Harpalus cardoni
Harpalus caspius
Harpalus cautus
Harpalus chalcopterus
Harpalus chasanensis
Harpalus chengjiangensis
Harpalus chiloschizontus
Harpalus chobautianus
Harpalus chrysopus
Harpalus cilihumerus
Harpalus circumpunctatus
Harpalus cisteloides
Harpalus cohni
Harpalus colchicus
Harpalus compar
Harpalus compressus
Harpalus contemptus
Harpalus convexus
Harpalus cordatus
Harpalus cordifer
Harpalus coreanus
Harpalus corporosus
Harpalus corpulentus
Harpalus corrugatus
Harpalus crates
Harpalus cupreus
Harpalus cursorius
Harpalus cyanopterus
Harpalus cyclogonus
Harpalus cyrtonotoides
Harpalus danieli
Harpalus davidi
Harpalus davidianus
Harpalus decipiens
Harpalus defector
Harpalus demelti
432 additional siblings truncated for brevity.
See the resource file for a full list.

wikidata_hierarchy tar gz

Biota
Eukaryota
Unikonta
Opisthokonta
Holozoa
Filozoa
Apoikozoa
Animalia
Eumetazoa
Bilateria
Protostomia
Ecdysozoa
Arthropoda
Hexapoda
Insecta
Dicondylia
Pterygota
Metapterygota
Neoptera
Eumetabola
Endopterygota
Coleoptera
Adephaga
Caraboidea
Carabidae
Harpalinae
Harpalus
Harpalus pennsylvanicus (this page)
Harpalus actiosus
Harpalus acupalpoides
Harpalus adenticulatus
Harpalus aeneipennis
Harpalus aenigma
Harpalus aequicollis
Harpalus aesculanus
Harpalus agakhaniantzi
Harpalus agilis
Harpalus akinini
Harpalus alajensis
Harpalus albanicus
Harpalus alexandrae
Harpalus alexeevi
Harpalus alienus
Harpalus alpivagus
Harpalus amarellus
Harpalus amariformis
Harpalus amplicollis
Harpalus amputatus
Harpalus anatolicus
Harpalus angulatus
Harpalus angustipennis
Harpalus angustitarsis
Harpalus animosus
Harpalus anisodactyliformis
Harpalus anxioides
Harpalus apache
Harpalus araraticus
Harpalus arcuatus
Harpalus arnoldii
Harpalus ascetes
Harpalus asemus
Harpalus asphaltinus
Harpalus aterrimus
Harpalus atratus
Harpalus atrichatus
Harpalus atripes
Harpalus attenuatus
Harpalus australasiae
Harpalus autumnalis
Harpalus azumai
Harpalus babai
Harpalus badakschanus
Harpalus baleensis
Harpalus balli
Harpalus basanicus
Harpalus basilewskyi
Harpalus basuto
Harpalus bellieri
Harpalus beneshi
Harpalus bitinctus
Harpalus brachypterus
Harpalus brevicornis
Harpalus brevis
Harpalus breviusculus
Harpalus brunnipes
Harpalus bucharicus
Harpalus bungii
Harpalus caeruleatus
Harpalus caiphus
Harpalus calathoides
Harpalus calceatus
Harpalus capicola
Harpalus cardoni
Harpalus caspius
Harpalus cautus
Harpalus chalcopterus
Harpalus chasanensis
Harpalus chengjiangensis
Harpalus chiloschizontus
Harpalus chobautianus
Harpalus chrysopus
Harpalus cilihumerus
Harpalus circumpunctatus
Harpalus cisteloides
Harpalus cohni
Harpalus colchicus
Harpalus compar
Harpalus compressus
Harpalus contemptus
Harpalus convexus
Harpalus cordatus
Harpalus cordifer
Harpalus coreanus
Harpalus corporosus
Harpalus corpulentus
Harpalus corrugatus
Harpalus crates
Harpalus cupreus
Harpalus cursorius
Harpalus cyanopterus
Harpalus cyclogonus
Harpalus cyrtonotoides
Harpalus danieli
Harpalus davidi
Harpalus davidianus
Harpalus decipiens
Harpalus defector
Harpalus demelti
432 additional siblings truncated for brevity.
See the resource file for a full list.
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