Foxiphalus secasius Barnard & Barnard 1982

Foxiphalus secasius

Paraphoxus obtusidens.—J.L. Barnard, 1960:249–259, pls. 33–37 [in part, especially pl. 36: fig. p (= AHF 456); not other parts and Alderman, 1936].

DESCRIPTION OF HOLOTYPE (female “f”).—Head about 21 percent of total body length, greatest width about 65 percent of length, rostrum unconstricted, broad, elongate, almost reaching apex of article 2 on antenna 1. Eyes medium, clear of pigment, ommatidia especially small. Article 1 of peduncle on antenna 1 about 1.7 times as long as wide, about twice as wide as article 2, ventral margin with about 8 setules, weakly produced dorsal apex with 2 setules-setae, article 2 about 0.45 times as long as article 1, with ventral tightly constrained double crescent of 8 setae, and 5 apicofacial setae, primary flagellum with 14 articles, about 1.1 times as long as peduncle, lacking aesthetascs (possibly abraded), accessory flagellum with 13 articles. Antenna 2 not ensiform; spine formula of article 4 = 1-3-4-5, dorsal margin with notch bearing 2 setae and 1 spine, ventral margin with 5 groups of 2–5 long to medium setae, 1 ventrodistal long spine set facially, article 5 about 0.8 times as long as article 4, facial spine formula = 3-3, dorsal margin bearing 3 small apical setae, ventral margin with 4 sets of 2–4 long to short setae, 3 ventrodistal long to medium spines, 1 of these facial; flagellum about 1.6 times as long as article 4–5 of peduncle combined, with 16 articles.

Epistome broadly rounded anteriorly; upper lip ordinary and dominant. Mandibles with medium palpar hump, right incisor with 3 teeth, left incisor with 3 humps in 2 branches; right lacinia mobilis bifid, distal branch much shorter than proximal, both branches simple, pointed, left lacinia mobilis with 4 teeth plus 2 accessory teeth, middle teeth shortened, right rakers 8, plus 1 rudimentary, left rakers 10 plus 1 rudimentary; molars composed of elongate plaques, right molar with 7 primarily long spines plus 1 short spine strongly disjunct, left molar with 6 primarily long spines plus 1 short spine strongly disjunct, each molar with plume; palp article 1 short, article 2 with 1 short inner apical seta and 2 other short inner setae, article 3 about 0.9 times as long as article 2, oblique apex with 9 spine-setae, basofacial formula = 0–2. Each outer lobe of lower lip with cone. Inner plate of maxilla 1 large, broad, bearing 1 long apical pluseta, 1 shorter apicomedial seta, 2 apicolateral much shorter setae, outer plate with 11 spines, palp article 2 with 1 apical spine, 1 apicolateral, 5 medial and 1 submarginal setae. Plates of maxilla 2 extending subequally, outer not broader than inner, outer with 5 apicolateral setae, inner with 2 medial setae. Inner plate of maxilliped with 1 large thick apical spine, 2 apicofacial setae, 3 medial setae, outer plate with 8 medial spines, 5 lateral setae, no cusp, palp article 1 without apicolateral seta, article 2 with 1 apicolateral seta, medial margin of article 2 moderately setose, article 3 with 5 and 3 facial setae in 2 continuous rows, 5 lateral setae in 2 groups, nail of article 4 fused to article, with 2 accessory setules.

Coxa 1 strongly expanded, anterior margin almost straight, main ventral setae of coxae 1–4 = 7-6-6-3, posteriormost seta of coxae 1–3 shortest or shortened, anterior and posterior margins of coxa 4 strongly divergent, posterior margin oblique, almost straight, posterodorsal corner rounded, posterodorsal margin ordinary, concave, width-length ratio of coxa 4 = 1:1. Gnathopods generally ordinary, width ratios on articles 5–6 of gnathopods 1–2 = 24:36 and 25:38, length ratios = 65:66 and 52:63, palmar humps large, palms oblique, article 5 of gnathopod 1 elongate, ovate, posterior margin flat, long, article 5 of gnathopod 2 short, posterior margin rounded, short, weakly lobate.

Pereopod 4 slightly stouter than pereopod 3 especially on article 4, facial setae formula on article 4 = 3 and 3, parallel to apex, on article 5 = 4 and 5, main spine of article 5 extending to M. 80 on article 6, article 5 with 2 and 1 proximoposterior spines, spine formula of article 6 = 4 + 5 and 4 + 5 plus small middistal seta, some spines especially long, acclivity on inner margin of dactyls of pereopods 3–4 weak, emergent setule almost fully immersed, midfacial pluseta ordinary. Coxae 5–7 posteroventral seta formula = 7-7-7; articles 4–5 of pereopods 5–6 narrow, facial spine rows sparse to moderately developed, facial ridge formula of article 2 on pereopods 5–7 = 0-1-1; width ratios of articles 2, 4, 5, 6 of pereopod 5 = 45:31:30:15, of pereopod 6 = 76:37:25:13, of pereopod 7 = 85:21:20:10, length ratios of pereopod 5 = 80:33:47:55, of pereopod 6 = 96:66:56:87, of pereopod 7 = 100:30:32:41, article 2 of pereopod 7 reaching middle of article 4, posterior margin with 10 small serrations, medial apex of article 6 scarcely combed, bearing 3–4 weak digital processes.

Posteroventral corner of epimeron 1 rounded, posterior margin straight, weakly serrate and with 3 setules, corner with setule, anteroventral margin with 11 medium setae, posteroventral face with 5 long setae set in vertical row. Posteroventral corner of epimeron 2 rounded, posterior margin straight, weakly serrate, with 3 setules, facial setae = 7, posteriormost triad set vertically. Posteroventral corner of epimeron 3 rounded, weakly protuberant, with tiny sinus, posterior margin straight but oblique, serrate, setose (11), ventral margin with 1 seta near middle.

Urosomite 1 with lateral setule at base of uropod 1, articulation line almost complete, urosomite 3 unprotuberant dorsally. Rami of uropods 1–2 with articulate but tightly fixed apical nails, except inner ramus of uropod 1 with flexible nail, outer ramus of uropod 1 with 4 tightly constrained dorsal spines, inner with 2, outer ramus of uropod 2 with 3 dorsal spines, inner with 1 dorsomedial spine. Peduncle of uropod 1 with 5 short basofacial setae, and 3 apicolateral spines, medially with 4 marginal setae and spines, apicalmost an ordinary spine, no displaced spine. Peduncle of uropod 2 with 8 short stout dorsal spines, medially with 1 small apical spine; apicolateral corners of peduncles on uropods 1–2 without comb. Peduncle of uropod 3 with 5 ventral spines, dorsally with 1 lateral spine and 2 setules, 1 medial spine and 1 setule, rami submasculine, inner extending to M. 95 on article 1 of outer ramus, apex with 3 setae, medial and lateral margins setose apically, article 2 of outer ramus short, 0.12, bearing 2 long setae, medial margin of article 1 setose, lateral margin with 2 acclivities, spine formula = 2-2-2, setal formula = 0-0-1. Telson especially long, length-width ratio = 17:14, almost fully cleft, each apex wide, rounded, lateral acclivity deep, with small lateral and medial spines separated by elongate setule, mid-lateral setules diverse, largest medium.

HOLOTYPE.—AHF 3512, female “f,” 8.61 mm.

TYPE-LOCALITY.—Velero III 456–35, Panama, Islas Secas, 07°57′50″N, 82°01′15″W, 12 fms (22 m), bottom of sand and nullipores, 6 Feb 1935.

MATERIAL.—Velero III 449–35 (1 female).

RELATIONSHIP.—Differing from the F. obtusidens group in the absence of a displaced peduncular spine on uropod 1; like F. apache but with rounded anterior margin of epistome, only 1 ventral seta on epimeron 3, and vertically arranged setae on epimeron 2.

See F. golfensis.

DISTRIBUTION.—Panama, Secas Islands, 12–25 fms (22–46 m).

Eobrolgus J.L. Barnard

Eobrolgus J.L. Barnard, 1979b:376.—Barnard and Barnard, 1981:295.

DIAGNOSIS.—See J.L. Barnard, 1979b.

TYPE-SPECIES.—Paraphoxus spinosus Holmes, 1905, original designation.

COMPOSITION.—Eobrolgus spinosus, E. chumashi.