Endomychidae, or handsome fungus beetles, is a family of beetles with representatives found in all ecozones. There are around 120 genera and 1300 species. The family was established based on the type genus Endomychus, a genus erected in 1795 by Panzer which was applied to a species that Linnaeus called as Chrysomela coccinea. As the common name suggests, Endomychidae feed on fungi. Crowson, in his influential treatment of the beetles, placed the family within the Cucujoidea. They have a tarsal formal of 4-4-4 or 3-3-3 and the wings lack a closed radial cell. The second antennal segment has a sensory appendage that is as long as the third antennal segment. The family has also been grouped with the Coccinellidae in a group called the Trimera for having pseudotrimerous tarsi. A 2015 molecular phylogeny study found that the Cucujoidea were found to be non-monophyletic and the Endomychidae was refined with the removal of the Anamorphinae from within the family and elevated to the status of a full family. Mycetaeinae and Eupsilobiinae were also found not to belong within the clades of the core Endomychidae.
The subfamilies that are included:Danascelinae Endomychinae (including Stenotarsinae) Epipocinae Leiestinae Lycoperdininae Merophysiinae Pleganophorinae Xenomycetinae