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BiologyHigh-oceanic, nyctoepipelagic at the surface and to 950 m and generally found between 475-850 m during the day (Ref. 4479). Neustonic to mesopelagic (Ref. 58302). Oviparous, with planktonic eggs and larvae (Ref. 31442). Lipid content is 3.8 % in fresh body weight and wax ester is 10.6 % in total lipids (Ref. 9193). Females reach sexual maturity at 4.8 cm, males at 3.5 cm (Ref. 47377).