Overview

Distribution

Distribution: China, India, Korea, Japan, Caucasus, Russia, Europe, North, Central, and South America.
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Physical Description

Morphology

Description

Plants moderate-sized, weak-stemmed, lax; yellowish to golden brown; forming loose carpets; branches loosely imbricate. Stems yellow to brown, superficial cortical layer with spiral reinforcing fibrils visible, 1 or more pores/cell, comb-fibrils on interior wall. Stem leaves short-rectangular, 0.8-1.1 mm, hyaline cells mostly non-septate and absent comb-fibrils. Branch fascicles with 2 spreading and 1-2 hanging branches. Branch stems with hyaline cells non-ornamented, no or weak funnel-like projections on the end walls of cortical cells, cortical cell walls usually with large round pores. Branch leaves ovate to ovate-elliptic, 1.4-1.8 mm; hyaline cells on convex surface with numerous pores along the commissures; comb-lamellae obvious only in proximal 1/2 of leaf; chlorophyllous cells broadly triangular in transverse section and well-enclosed on the convex surface. Sexual condition dioecious. Spores (22) 24-27(-28) µm, surface granulate.
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Description

Plants 7–15 cm high, green to yellowish green or yellowish brown, in loose or extensive, compact cushions. Stem cortex in 3–5 layers, hyaline cells large and thin-walled, with fibrils and pores; central cylinder yellowish brown. Stem leaves 1.0–1.8 mm × 0.8–1.0 mm, short, broadly ligulate, slightly narrowed at base, flat in the upper part, lacerate at the rounded apex; hyaline cells generally divided, without fibrils, or rarely with indistinct fibrils. Branches in fascicles of 4–5, with 2 spreading. Branch leaves 1.5–2.5 mm long, imbricate, concave, broadly ovate to rounded ovate, cucullate-concave near the apex, hyaline and slightly dentate at the apex, marginal border indistinct; hyaline cells densely fibrillose, with numerous commissural pores on the dorsal surface, with pores at the upper cell angles becoming larger and fewer near the bases on the ventral surface, the inner walls adjacent to green cells often dense with fringe-fibrils; green cells in cross section isosceles-triangular, exposed on both surfaces, more broadly so on the ventral surface. Dioicous; antheridial branches reddish brown. Perigonial leaves large, ovate; perichaetia growing at base of capitulum branches. Spores yellowish brown, smooth.
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Ecology

Habitat

Habitat: in bogs or wet humic soil under coniferous forests.
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Notes

Comments

Sphagnum imbricatum to be East Asian in distribution but a recent collection in Selawik National Wildlife Refuge places it in the North American flora. It will undoubtedly be found elsewhere. Sphagnum imbricatum is closest in morphological detail to S. steerei, but the latter is a very dark colored and densely branched species whereas S. imbricatum is quite light in color and not particularly dense.
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© Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA

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