Overview
Brief Summary
Biology
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Comprehensive Description
Description
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Distribution
Global Range: (>2,500,000 square km (greater than 1,000,000 square miles)) Breeds from central and southeastern Alaska east across northern Canada and southern Victoria Island to western Melville Peninsula, northeastern Manitoba, northern Ontario, northern Quebec, and southern Baffin Island (recently naturally established in western Greenland) south to southwestern British Columbia, northeastern California, northern Utah, south-central Wyoming, South Dakota, Kansas, northern Arkansas, western Kentucky, southern Ohio, Pennsylvania, northern Virginia, and Maryland. Breeding populations in the southern prairie states were extirpated, but many have become reestablished. Birds in eastern states south of Great Lakes and Massachusetts result from relatively recent natural southward extension of breeding range and to great extent from introductions. Feral populations resulting from introductions may occur almost anywhere in the United States (AOU 2004).
Winters from the southern part of the breeding range through most of the United States and into northern Mexico. Introduced and established in Great Britain, Iceland, southern Scandinavia, and New Zealand. Accidental in Hawaii, Greater Antilles, and the Bahamas.
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Geographic Range
Canada Geese are found throughout North America. There are four subspecies (or populations); each is found in a different area of North America. These subspecies are the southern, northern, western and Aleutian-Canadian populations. The southern population ranges from 60 degrees north latitude to the Rockies and Atlantic Ocean. The northern population ranges north of 60 degrees north latitude in the Arctic and Subarctic. Canada geese travel to the southern parts of the United States during the winter. The western population is found along the coasts of Alaska and British Columbia. The Aleutian-Canadian population is rarely found. A general trend in all subspecies is that they they spend summers in the northern parts of North America, especially Canada, and migrate south to areas of the United States in the winter months.
Biogeographic Regions: nearctic (Native )
- Ogilvie, M. 1978. Wild Geese. Vermillion, SD: Buteo Books.
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Range Description
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National Distribution
Canada
Origin: Native
Regularity: Regularly occurring
Currently: Present
Confidence: Confident
Type of Residency: Year-round
United States
Origin: Native
Regularity: Regularly occurring
Currently: Present
Confidence: Confident
Type of Residency: Year-round
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Range
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Physical Description
Morphology
Physical Description
Branta canadensis individuals have a black neck, bill, and head with a white strap under the chin and occasional white patches elsewhere. The body is usually brownish-gray although colors vary in some of the subspecies. In some of the smaller subspecies the body is dark brown in color where as in some of the larger subspecies, the body is a light gray tone. Underneath, the colors are much lighter and almost white on the tail. During flight the tail shows a white semi-circle just above the black tail. Females may be slightly smaller than males, although both are similar to each other in color pattern. The bill of Branta canadensis tapers from the base where it is high to the end where it has narrowed. The bill has lamellae, or teeth around the outside that are a used as a cutting tool. The legs are close together with very black feet. These geese have very large wings (127 to 173 cm wingspan) that can also be used as weapons. The weight of Branta canadensis varies depending on the subspecies.
Goslings (young Canada geese) are yellow with some greenish-gray colorings on top of their heads and backs. As with the adult color pattern, there is some variation among the different subspecies. Goslings of the darker subspecies have a brownish olive or blunt yellow coloring while those of the lighter subspecies are lighter and brighter in color. These colors fade as the gosling grows into the adult color pattern. All goslings have black or blue-gray bills and legs that become darker as they age.
Range mass: 1100 to 8000 g.
Range length: 90 to 200 cm.
Range wingspan: 127 to 173 cm.
Other Physical Features: endothermic ; homoiothermic; bilateral symmetry
Sexual Dimorphism: male larger
- Robbins, C., B. Bruun, H. Zim. 2001. A guide to field identification: Birds of North America. New York: St. Martin's Press.
- Kortright, F. 1942. The Ducks, Geese and Swans of North America. Washington, DC: The American Wildlife Institute.
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Size
Diagnostic Description
Differs from the brant (Branta bernicla) in having a broad white chin strap rather than a small whitish patch on either side of the neck. Differs from the barnacle goose (Branta leucopsis) by the lack of a mostly white face and by having plain dark wings instead of blue-gray upperparts barred with black (NGS 1983).
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Ecology
Habitat
Habitat
Canada Geese are found near waterways in open, grassy habitats such as grasslands, chaparral, and arctic tundra. They also inhabit man-made habitats that are open and grassy, such as golf courses, agricultural land, airports, and parks.
Habitat Regions: temperate ; terrestrial ; freshwater
Terrestrial Biomes: tundra ; savanna or grassland
Aquatic Biomes: lakes and ponds; rivers and streams; coastal
Wetlands: marsh
Other Habitat Features: suburban ; agricultural ; riparian
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Water temperature and chemistry ranges based on 2 samples.
Environmental ranges
Depth range (m): 0 - 0
Temperature range (°C): 5.940 - 13.008
Nitrate (umol/L): 2.446 - 3.533
Salinity (PPS): 30.572 - 33.176
Oxygen (ml/l): 6.169 - 7.377
Phosphate (umol/l): 0.377 - 0.659
Silicate (umol/l): 1.811 - 6.162
Graphical representation
Temperature range (°C): 5.940 - 13.008
Nitrate (umol/L): 2.446 - 3.533
Salinity (PPS): 30.572 - 33.176
Oxygen (ml/l): 6.169 - 7.377
Phosphate (umol/l): 0.377 - 0.659
Silicate (umol/l): 1.811 - 6.162
Note: this information has not been validated. Check this *note*. Your feedback is most welcome.
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Comments: Various habitats near water, from temperate regions to tundra. In migration and winter, coastal and freshwater marshes, lakes, rivers, fields, etc. On Admiralty Island, Alaska, commonly perches in trees. In the eastern U.S., common on lawns adjacent to water in urban-suburban areas.
Breeds in open or forested areas near lakes, ponds, large streams, inland and coastal marshes. The nest is built on the ground or on an elevated place (muskrat house, abandoned heron's nest, rocky cliffs, etc.) (Terres 1980). Usually returns to nesting territory used in previous year.
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Habitat
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Migration
Non-Migrant: Yes. At least some populations of this species do not make significant seasonal migrations. Juvenile dispersal is not considered a migration.
Locally Migrant: Yes. At least some populations of this species make local extended movements (generally less than 200 km) at particular times of the year (e.g., to breeding or wintering grounds, to hibernation sites).
Locally Migrant: Yes. At least some populations of this species make annual migrations of over 200 km.
Found year-round in central part of range. Flocks of family groups migrate together.
The percentage of the population wintering in the north is now higher than in the past, due at least in part to increased availability of planted corn. A large percentage of geese that spend a winter in the Carolinas winter in more northerly locations (e.g., Chesapeake) in subsequent winters (Ecology 72:523).
Some western geese, mainly prebreeders and unsuccessful nesters, make molt migrations to and from molting areas during and after the brood-rearing season; apparently, molters of Pacific and Rocky Mountain populations that leave those regions go to the Northwest Territories of Canada (Krohn and Bizeau 1980). See Johnson and Herter (1989) for information on migration of various subspecies.
In the Mississippi Valley population, fall and spring migration is concentrated in a corridor in central Illinois and southeastern Wisconsin; a major spring staging area is the Kaskaskia River Valley in central Illinois; chronology of departure from wintering areas and of fall migration vary annually (Tacha et al. 1991). See Samuel et al. (1991) for information on fall and winter distribution in the Mississippi Flyway.
Atlantic Flyway population consists of migrant geese and a resident population which generally migrates only short distances; migrant population breeds primarily in Labrador, northern Quebec, and James Bay area and winters south to North Carolina, western Pennsylvania, the western Carolinas, and Mississippi Flyway states.
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Trophic Strategy
Food Habits
When on land, Branta canadensis eat a variety of grasses including Bermuda grass, salt grass and wild barley. Geese are able to grab a hold of each blade and pull it out with their bills by jerking their heads. They also eat wheat, beans, rice, and corn. In the water, the birds stick their head and upper part of their body into the water leaving their tail and back end extending in the air. They stretch their neck out, under the water, and slide their bills across the bottom silt. They also eat a number of aquatic plants such as eel grass, sea lettuce and sago.
Plant Foods: leaves; roots and tubers; seeds, grains, and nuts; algae
Primary Diet: herbivore (Folivore )
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Comments: Grazes on marsh grasses, sprouts of winter wheat (spring), grain (fall); eats clover, cattails, bulrushes, algae, pond- weed, and other plants. Feeds in shallows, marshes, fields. Also eats mollusks and small crustaceans (Terres 1980). Subspecies OCCIDENTALIS of west coastal North America: exogenous sources of lipid and protein are important to energy and nutrient requirements of nesting geese (Condor 95:193-210).
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Associations
Ecosystem Roles
As well as dispersing the seeds of the plants they eat, Canada Geese are important prey for many predators in the ecosystems in which they live.
Ecosystem Impact: disperses seeds
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Predation
Unguarded nests and eggs are targets for predators such as gulls, common ravens, American crows, skunks, domestic dogs, and many others. Males send out an alarm by flying into the air and honking as a predator approaches. This alerts not only his mate but others nesting nearby. Females lower their bodies onto the nest and stretch out their necks to camouflage the nest.
Canada geese are also a common game bird, hunted regularly by humans.
Known Predators:
- gulls (Laridae)
- common ravens (Corvus corax)
- American crows (Corvus brachyrhynchos)
- striped skunks (Mephitis mephitis)
- domestic dogs (Canis lupus familiaris)
- humans (Homo sapiens)
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Known predators
Laridae
Corvus corax
Corvus brachyrhynchos
Homo sapiens
Mephitis mephitis
Canis lupus familiaris
This list may not be complete but is based on published studies.
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Known prey organisms
algae
Larus californicus
This list may not be complete but is based on published studies.
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Population Biology
General Ecology
Mean annual survival rate for Rocky Mountain birds banded on nesting areas: 53% for immatures, 64% for adults (Krohn and Bizeau 1980).
In winter, flocks foraged up to 48 km from roost in Texas (Glazener 1946).
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Life History and Behavior
Behavior
Communication and Perception
Canada geese have good eyesight, which is necessary for flight. They must move their heads in order to see all the way around themselves. However, their eyes are close to their crowns on the side of their heads, enabling them to see more than 180 degrees (closer to 270 degrees) horizontally and vertically. They have mostly monocular vision. Canada geese have excellent hearing and the ears are located on the side of its head. Canada geese often use body movements to communicate with each other. These geese also have the ability to make at least 10 different calls
Communication Channels: visual ; tactile ; acoustic
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Cyclicity
Comments: May be active day or night during migration. Usually feeds in early morning and late afternoon.
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Life Expectancy
Lifespan/Longevity
It is not clear exactly how long the average Canada goose can live, but there have been two geese that were reported to have lived very long lives. One of them lived to be 24 years old and another reached 23. In captivity, two geese were reported to live to 42 years old. Probably most Canada geese die within their first year of life, as nestlings, fledglings, and during their first migration.
Range lifespan
Status: wild: 24 (high) years.
Range lifespan
Status: captivity: 42 (high) years.
Average lifespan
Status: wild: 364 months.
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Lifespan, longevity, and ageing
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Reproduction
Reproduction
Canada geese are monogamous. Pairs form during the winter, during migration or on their wintering grounds, for the next breeding season. Mated pairs may stay together for more than one year, sometimes staying together for life.
Males fight over females with their wings and bills. The winner approaches the female with his head down and neck undulating. He makes hissing and honking noises. The pairs mate either before or after they have found a nesting location. Mating, occurs in the spring on the water. The female is usually partially submerged or completed submerged while copulation takes place.
Mating System: monogamous
The average clutch size is five eggs, although this size ranges from 2 to 9 eggs. The incubation period lasts 23 to 30 days.
Females incubate the eggs, choose the location for nesting, and even build the nest without males. Males defend the territory, nest, and eggs from intruders, such as other geese. Female Canada geese pick nesting sites that are isolated but have good visibility. This allows them to readily see danger approaching and to be difficult to get at. The nesting area also must have open water with low banks so they can have access to water plants and places to get into or out of the water. Swamps, marshes, meadows, lakes, and other such areas are among some of their favorite nesting spots. The Canadian and Alaskan shorelines have expanses of tundra habitat that provide good nesting sites. Canada geese are often seen nesting on small islands that don't have very tall grasses or on muskrat houses (which are similar to small islands).
Nests are very simple and are made quickly. Materials that are used are weeds, twigs, grass, moss, needles, and other such materials. After some collection and building, female geese round out a curve or depression with their bodies. They drop the materials around themselves and move the items to get the best fit. From time to time they round out the center with their chests or feet. If there are no items of vegetation the nest may only be a depression in the ground shaped by their chests and feet. Once the eggs are laid, the nest is lined with feathers and down. Down insulates against extreme warmth as well as cold, stabilizing egg temperature.
Incubation must occur immediately after the last eggs are laid. The incubation period lasts 23 to 30 days. The female turns the eggs regularly to promote proper development and changes their position in the nest to maintain even incubation temperatures. The offspring hatches via an egg tooth on top of its beak to crack open the shell. Goslings keep cracking open the shells until they are completely free between 24 and 48 hours later. All of the eggs in the clutch are fully hatched within 24 hours. Goslings within a clutch usually have a sex ratio of 1:1. Hatchlings fledge in 68 to 78 days after hatching.
Breeding interval: Canada geese breed once yearly.
Breeding season: Females start laying eggs during the first weeks of March and continue as late as June in parts of the Arctic.
Range eggs per season: 2 to 9.
Average eggs per season: 5.
Range time to hatching: 23 to 30 days.
Range fledging age: 68 to 78 days.
Average age at sexual or reproductive maturity (female): 1 years.
Average age at sexual or reproductive maturity (male): 1 years.
Key Reproductive Features: seasonal breeding ; gonochoric/gonochoristic/dioecious (sexes separate)
Average eggs per season: 5.
After the eggs hatch, the family group (the offspring and parents) leave the nest and begin to travel together to feed and seek shelter. Both males and females feed and guard their young. Upon hatching, young Canada geese are able to follow their parents around and leave the nest.
Parental Investment: precocial ; pre-fertilization (Provisioning, Protecting: Female); pre-hatching/birth (Provisioning: Female, Protecting: Female); pre-weaning/fledging (Provisioning: Male, Female, Protecting: Male, Female); pre-independence (Provisioning: Male, Female, Protecting: Male, Female)
- Ehrlich, P., D. Dobkin, D. Wheye. 1988. The Birder's Handbook: A Field Guide to the Natural History of North American Birds. New York: Simon and Schuster.
- 1998. "Canada Geese (*Branta canadensis*)" (On-line). Accessed March 12, 2000 at http://www.rhrwildlife.com/cgee.htm.
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Clutch size 2-11 (usually 5-6). Incubation 25-30 days, by female (Harrison 1978). Nestlings precocial. Young tended by both adults, remain with adults until next spring. Some individuals begin breeding at 2 years, most by age 3 years.
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Evolution and Systematics
Functional Adaptations
Functional adaptation
Down feathers of geese insulate through special architecture.
"Feather keratin occurs in a 'b-sheet' configuration which differs from the a-helices that occur in mammalian keratins. . . We have measured the properties of individual down feathers from ducks, geese and penguins and found that their properties are similar to flight feathers and, indeed, the man-made polymers used in artificial insulation fibres. The message is that the architecture of down feathers is probably more important than material properties in determining their advantages over synthetic materials. . .Recently, we have begun to explore the toughness of feather keratin by using instrumented clippers and scissors. The fracture toughness of β-keratin has proved to be very high, around 10 kJ m-2." (Bonser 2007)
Learn more about this functional adaptation.
- Cameron, G. J.; Wess, T. J.; Bonser, R. H. C. 2003. Young’s modulus varies with differential orientation of keratin in feathers. Journal of Structural Biology. 143(2): 118-123.
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Bonser, Richard. 2008. Mechanical properties of keratin.
http://www.rdg.ac.uk/biomim/personal/richard/keratin.htm.
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Molecular Biology and Genetics
Molecular Biology
Barcode data: Branta canadensis
There are 129 barcode sequences available from BOLD and GenBank. Below is a sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species. See the BOLD taxonomy browser for more complete information about this specimen and other sequences.
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Download FASTA File
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Statistics of barcoding coverage: Branta canadensis
Public Records: 129
Species: 425
Species With Barcodes: 1
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Conservation
Conservation Status
IUCN Red List Assessment
Red List Category
Red List Criteria
Version
Year Assessed
Assessor/s
Reviewer/s
Contributor/s
Justification
History
- 2008Least Concern
- 2006Least Concern
- 2004Not Recognized
- 2000Not Recognized
- 1994Not Recognized
- 1988Not Recognized
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Conservation Status
In 1918 when the Migratory Bird Treaty was passed, spring shooting was prohibited in the United States and Canada. This regulated the hunting season to three and a half months of the year. The hunting regulations currently in place are for shooting season limits and bag limits in relation to the amount of birds currently in the population. A quota system was put in place in 1960 to regulate the number of geese shot in a given year.
One subspecies, Aleutian-Canadian Geese, were listed as endangered by the U.S. Fish and Wildlife Service in 1967. This was due to the introduction of a non-native arctic fox species to their nesting islands. They became predatory on the naturally defenseless geese. This introduction caused the population to decline to approximately 800 individuals. However, in 1990, due to increasing numbers, the U.S. Fish and Wildlife Service changed the listing to threatened. The state of Alaska also changed the species listing from endangered to a species of special concern. Aleutian-Canadian Geese are now recorded around 15,000 individuals and nesting on eight islands.
On the other hand, some populations have grown so numerous, there are many organizations who are trying to regulate the populations of these geese. They see this as necessary because if the goose population continues to rise at its current rate, they will present a very serious problem to their surrounding environment in only a few years. Other organizations believe that the methods and ideas of these organizations are cruel and unnecessary. These groups believe that the growing population is not nearly as threatening as some believe and that they are actually at great risk because of the excessive hunting and death by pesticides that geese populations experience.
US Migratory Bird Act: protected
US Federal List: threatened
CITES: no special status
State of Michigan List: no special status
IUCN Red List of Threatened Species: least concern
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National NatureServe Conservation Status
Canada
Rounded National Status Rank: N5B,N5N : N5B: Secure - Breeding, N5N: Secure - Nonbreeding
United States
Rounded National Status Rank: N5B,N5N : N5B: Secure - Breeding, N5N: Secure - Nonbreeding
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Status
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Trends
Threats
Threats
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Comments: Pesticide use is a potential threat; for example, 1600 died in a wheat field that had been sprayed with parathion (see Franson 1994).
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Management
Management Requirements: Methods for reducing goose problems on lawns include habitat modification (such as planting vegetation that reduces long-distance visibility or that interferes with take-offs or landings, draining bodies of water, or eliminating palatable vegetation), use of fear-provoking stimuli, and hunting (see Converse 1985, Conover and Chasko 1985, Conover 1991, and Conover and Kania 1991 for further information on resident nuisance geese in the northeastern U.S.). See Conover (1989) for information on the use of methiocarb to reduce damage to grain fields and winter cover crops. See Cummings et al. (1991) for information on the use of dimethyl anthranilate and methyl anthranilate to repell Canada geese from grassy areas. In Wisconsin, farmers rated mylar flags and human effigies as fully successful or at least helpful in reducing crop damage; Av-Alarm (a sonic deterrent) also was effective but not well accepted by farmers (Heinrich and Craven 1990). See Aguilar et al. (1991) for an evaluation of goose alarm or distress calls and screamer shells to disperse wintering urban geese.
See Reese et al. (1987) for recommendations on construction of artificial nesting islands.
Tacha et al. (1991) found no evidence to support the existence of manageable subpopulations in the Mississippi Valley population.
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Needs: Protect migratory population in Atlantic Flyway while reducing size of nuisance resident population (USFWS 1996).
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Conservation
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Relevance to Humans and Ecosystems
Benefits
Economic Importance for Humans: Negative
Canada geese can become a nuisance, especially when normally migratory birds become resident. They can overgraze lawns and crops, leading to erosion. On lawns, their feces can annoy humans. Build-up of fecal matter can lead to reduced water quality, by fostering bacteria and adding much nitrogen and phosphorus.
Canada geese can be an exceptional annoyance in Atlantic flyway states by crowding in on golf courses, beaches, parks, playing fields, and yards. In the eastern states, Canada Geese have been very harmful to local crops and have forced farmers to plant a lot more winter wheat to compensate for the damage done by these geese.
Negative Impacts: crop pest
- NJ Department of Environmental Protection, 2001. "Management of Canada geese in urban areas" (On-line). Accessed 14 May 2002 at http://www.state.nj.us/dep/watershedmgt/DOCS/BMP_DOCS/Goosedraft.pdf.
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Economic Importance for Humans: Positive
Canada Geese have been hunted by humans for hundreds of years. Native Americans hunted them in the spring migration. Eskimos hunted them by taking advantage of the molt that leaves them flightless. Even early white settlers took advantage of these birds and hunted them for food. These birds are still being hunted today in the United States and Canada.
Positive Impacts: food ; body parts are source of valuable material
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Economic Uses
Comments: Sport hunting accounts for more than 86% of the mortality of fledged geese in Rocky Mountain region (Krohn and Bizeau 1980).
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Wikipedia
Canada Goose
The Canada Goose (Branta canadensis) is a wild goose with a black head and neck, white patches on the face, and a brownish-gray body. Native to arctic and temperate regions of North America, it is occasionally found in northern Europe, and has been introduced to other temperate regions.
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Taxonomy and etymology
The Canada Goose was one of the many species described by Carl Linnaeus in his 18th-century work Systema Naturae.[2] It belongs to the Branta genus of geese, which contains species with largely black plumage, distinguishing them from the grey species of the Anser genus. The specific epithet canadensis is a New Latin word meaning "from Canada". According to the Oxford English Dictionary, the first citation for the 'Canada Goose' dates back to 1772. The Canada Goose is also referred to as the Canadian Goose,[3] although that name is discouraged.[4][5][verification needed]
The Cackling Goose was originally considered to be the same species or a subspecies of the Canada Goose, but in July 2004 the American Ornithologists' Union's Committee on Classification and Nomenclature split the two into two species, making Cackling Goose into a full species with the scientific name Branta hutchinsii. The British Ornithologists' Union followed suit in June 2005.[6]
The AOU has divided the many subspecies between the two animals. To the present species were assigned:
- Atlantic Canada Goose, Branta canadensis canadensis
- Interior Canada Goose, Branta canadensis interior
- Giant Canada Goose, Branta canadensis maxima
- Moffitt's Canada Goose, Branta canadensis moffitti
- Vancouver Canada Goose, Branta canadensis fulva
- Dusky Canada Goose, Branta canadensis occidentalis
- part of "Lesser complex", Branta canadensis parvipes
The distinctions between the two geese have led to confusion and debate among ornithologists. This has been aggravated by the overlap between the small types of Canada Goose and larger types of Cackling Goose. The old "Lesser Canada Goose" was believed to be a partly hybrid population, with the birds named taverneri considered a mixture of minima, occidentalis and parvipes. In addition, it has been determined that the Barnacle Goose is a derivative of the Cackling Goose lineage, whereas the Hawaiian Goose is an insular representative of the Canada Goose.
Description
The black head and neck with a white "chinstrap" distinguish the Canada Goose from all other goose species, with the exception of the Barnacle Goose, but the latter has a black breast, and also grey, rather than brownish, body plumage.[7] There are seven subspecies of this bird, of varying sizes and plumage details, but all are recognizable as Canada Geese. Some of the smaller races can be hard to distinguish from the newly-separated Cackling Goose.
This species ranges from 75 to 110 cm (30 to 43 in) in length and has a 127–185 cm (50–73 in) wingspan.[8] The male usually weighs 3.2–6.5 kg (7.1–14 lb), and can be very aggressive in defending territory. The female looks virtually identical but is slightly lighter at 2.5–5.5 kg (5.5–12 lb), generally 10% smaller than its male counterpart, and has a different honk. An exceptionally large male of the race B. c. maxima, the "Giant Canada Goose" (which rarely exceed 8 kilograms (18 lb)), weighed 10.9 kilograms (24 lb) and had a wingspan of 2.24 metres (7.3 ft). This specimen is the largest wild goose ever recorded of any species. The life span in the wild of geese that survive to adulthood ranges 10–24 years.[9]
Distribution and habitat
This species is native to North America. It breeds in Canada and the northern United States in a variety of habitats. Its nest is usually located in an elevated area near water such as streams, lakes, ponds and sometimes on a beaver lodge. Its eggs are laid in a shallow depression lined with plant material and down. The Great Lakes region maintains a very large population of Canada Geese.[citation needed]
By the early 20th century, over-hunting and loss of habitat in the late 19th century and early 20th century had resulted in a serious decline in the numbers of this bird in its native range. The Giant Canada Goose subspecies was believed to be extinct in the 1950s until, in 1962, a small flock was discovered wintering in Rochester, Minnesota, by Harold Hanson of the Illinois Natural History Survey.[10] With improved game laws and habitat recreation and preservation programs, their populations have recovered in most of their range, although some local populations, especially of the subspecies occidentalis, may still be declining.[citation needed]
In recent years, Canada Goose populations in some areas have grown substantially, so much so that many consider them pests for their droppings, bacteria in their droppings, noise, and confrontational behavior. This problem is partially due to the removal of natural predators and an abundance of safe, man-made bodies of water near food sources, such as found on golf courses, in public parks and beaches, and in planned communities. Due in part to the interbreeding of various migratory subspecies with the introduced non-migratory Giant subspecies, Canada Geese are frequently a year-around feature of such urban environments.[citation needed]
Contrary to its normal migration routine, large flocks of Canada Geese have established permanent residence in Esquimalt, British Columbia, on Chesapeake Bay, in Virginia's James River regions, and in the Triangle area of North Carolina (Raleigh, Durham, Chapel Hill), and nearby Hillsborough. Some Canada Geese have taken up permanent residence as far south as Florida, in places such as retention ponds in apartment complexes. Some flocks never migrate, even during the winter, where food is available throughout the year.[citation needed]
Outside North America
Canada Geese have reached northern Europe naturally, as has been proved by ringing recoveries. The birds are of at least the subspecies parvipes, and possibly others. Canada Geese are also found naturally on the Kamchatka Peninsula in eastern Siberia, eastern China, and throughout Japan.[citation needed]
Greater Canada Geese have also been introduced in Europe, and have established populations in Great Britain, the Netherlands, Belgium, Germany, and Scandinavia. Semi-tame feral birds are common in parks, and have become a pest in some areas. In the early 17th century, explorer Samuel de Champlain sent several pairs of geese to France as a present for King Louis XIII. The geese were first introduced in Britain in the late 17th century as an addition to King James II's waterfowl collection in St. James's Park.[citation needed]
Canada Geese were introduced as a game bird into New Zealand and have also become a problem in some areas, fouling pastures and damaging crops. They were protected under the Wildlife Act 1953 and the population was managed by Fish and Game New Zealand who culled excessive bird numbers. In 2011 the government removed the protection status allowing anyone to kill the birds.[11] It was feared that farmers would resort to poisoning the birds although this was denied by Federated Farmers.[12]
Behavior
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Like most geese, the Canada Goose is naturally migratory with the wintering range being most of the United States. The calls overhead from large groups of Canada Geese flying in V-shaped formation signal the transitions into spring and autumn. In some areas, migration routes have changed due to changes in habitat and food sources. In mild climates from California to the Great Lakes, some of the population has become non-migratory due to adequate winter food supply and a lack of former predators.[citation needed]
Diet
Canada Geese are primarily herbivores,[13] although they sometimes eat small insects and fish.[14] Their diet includes green vegetation and grains. The Canada Goose eats a variety of grasses when on land. It feeds by grasping a blade of grass with the bill, then tearing it with a jerk of the head. The Canada Goose also eats grains such as wheat, beans, rice, and corn when they are available. In the water, it feeds from silt at the bottom of the body of water. It also feeds on aquatic plants, such as seaweeds.[9] In urban areas, they are also known to pick food out of garbage bins.
Reproduction
During the second year of their lives, Canada Geese find a mate. They are monogamous, and most couples stay together all of their lives.[9] If one dies, the other may find a new mate. The female lays 3–8 eggs and both parents protect the nest while the eggs incubate, but the female spends more time at the nest than the male.[citation needed]
Known egg predators include coyotes,[15] Arctic Foxes, Northern Raccoons, Red Foxes, large gulls, Common Raven, American Crows and bears.[16]
The incubation period, in which the female incubates while the male remains nearby, lasts for 24–28 days after laying. As the annual summer molt also takes place during the breeding season, the adults lose their flight feathers for 20–40 days, regaining flight at about the same time as their goslings start to fly.[17]
Adult geese are often seen leading their goslings in a line, usually with one parent at the front, and the other at the back. While protecting their goslings, parents often violently chase away nearby creatures, from small blackbirds to lone humans that approach, after warning them by giving off a hissing sound and will then attack with bites and slaps of the wings if the threat does not retreat or has seized a gosling. Most of the species that prey on eggs will also take a gosling. Although parents are hostile to unfamiliar geese, they may form groups of a number of goslings and a few adults, called crèches.[citation needed]
The offspring enter the fledging stage any time from 6 to 9 weeks of age. They do not leave their parents until after the spring migration, when they return to their birthplace. Once they reach adulthood, Canada Geese are rarely preyed on, but (beyond humans) can be taken by Coyotes, Red Foxes, Gray Wolves, Snowy Owls, Great Horned Owls, Golden Eagles and, most often, Bald Eagles.[16]
Relationship with humans
In North America, non-migratory Canada Goose populations have been on the rise. The species is frequently found on golf courses, parking lots and urban parks, which would have previously hosted only migratory geese on rare occasions. Owing to its adaptability to human-altered areas, it has become the most common waterfowl species in North America. In many areas, non-migratory Canada Geese are now regarded as pests by humans. They are suspected of being a cause of an increase in high fecal coliforms at beaches.[18] An extended hunting season, deploying noise makers, and hazing by dogs have been used in an attempt to disrupt suspect flocks.[19]
Since 1999, The United States Department of Agriculture Wildlife Services agency has been engaged in lethal culls of Canada Geese primarily in urban or densely populated areas. The agency responds to municipalities or private land owners, such as golf courses, who find the geese obtrusive or object to their waste.[20] Addling goose eggs and destroying nests are promoted as humane population control methods.[21]
Geese have a tendency to attack humans when they feel themselves or their goslings to be threatened. First the geese will stand erect, spread their wings and produce a hissing sound. Next, the geese will charge. They may then bite or attack with their wings.[22]
Canada Geese have been implicated in a number of bird strikes by aircraft; their large size and tendency to fly in flocks may exacerbate the effect. In 1995, a US Air Force E-3 Sentry aircraft at Elmendorf AFB, Alaska struck a flock of Canada Geese on takeoff and crashed, killing all 24 crew. The accident sparked efforts to avoid such events, including habitat modification, aversion tactics, herding and relocation, and culling of flocks.[23] A collision with a flock of migratory Canada Geese resulted in US Airways Flight 1549 suffering a total power loss after takeoff from LaGuardia Airport, New York City, New York on 15 January 2009. The pilot brought the plane to an emergency 'splash'-landing in the Hudson River, causing only minor injuries to the 155 passengers and crew.[24][25]
Migration
Canada Geese are known for their seasonal migrations. Most Canada Geese have staging or resting areas where they join up with others. Their autumn migration can be seen from September to the beginning of November. The early migrants have a tendency to spend less time at rest stops and go through the migration a lot faster. The later birds usually spend more time at rest stops. These geese are also renowned for their V-shaped flight formation. The front position is rotated since flying in front consumes the most energy. Canada Geese leave the winter grounds more quickly than the summer grounds. Elevated thyroid hormones, such as T3 and T4, have been measured in geese just after a big migration. This is believed because of the long days of flying in migration the thyroid gland sends out more T4 which will help the body cope with the longer journey. The increased T4 levels are also associated with increased muscle mass (hypertrophy) of the breast muscle, also because of the longer time spent flying. It is believed that the body sends out more T4 to help the goose's body with this long task by speeding up the metabolism and temperature at which the body works.[26] Also, other studies done show corticosterone levels to rise dramatically in these birds after and during a migration. Corticosterone is known a stress hormone, so it only makes sense that when these birds are stressed by flying long distances every day, that more corticosterone is released into their system. It is believed that a higher level of corticosterone will help the birds better manage this task.[27]
References
- ^ BirdLife International (2009). "Branta canadensis". IUCN Red List of Threatened Species. Version 2010.4. International Union for Conservation of Nature. http://www.iucnredlist.org/apps/redlist/details/141453. Retrieved 23 February 2011.
- ^ Linnaeus, C (1758) (in Latin). Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Holmiae. (Laurentii Salvii).
- ^ "Canadian goose". Dictionary.com. InterActiveCorp. http://dictionary.reference.com/browse/Canadian+goose. Retrieved 3 January 2011.
- ^ Gill, Frank, & Minturn Wright, Birds of the World: Recommended English Names Christopher Helm / Princeton University Press, 2006.
- ^ Gill, F, & D Donsker (eds), 2012. IOC World Bird Names (v. 3.1). Available at http://www.worldbirdnames.org/ Accessed 2012-05-01
- ^ Stackhouse, Mark. The New Goose.
- ^ Audubon Society
- ^ Ogilvie & Young, Wildfowl of the World. New Holland Publishers (2004), ISBN 978-1-84330-328-2
- ^ a b c Dewey, T.; Lutz, H. (2002). "Branta canadensis". Animal Diversity Web. http://animaldiversity.ummz.umich.edu/site/accounts/information/Branta_canadensis.html. Retrieved 2007-11-18.
- ^ Hanson, Harold C. (1997). The Giant Canada Goose (2nd. ed.). Southern Illinois University Press. ISBN 978-0-8093-1924-4.
- ^ "Canada geese protection status changed". Beehive - New Zealand Government. 17 March 2011. http://beehive.govt.nz/release/canada-geese-protection-status-changed. Retrieved 24 March 2011.
- ^ Cronshaw, Tim (25 March 2011). "Farmers deny poison plans". The Press.
- ^ Mowbray, Thomas B.; Ely, Craig R.; Sedinger, James S. and Trost, Robert E. (2002). "Canada Goose (Branta canadensis)". In Poole, A.. The Birds of North America. Ithaca: Cornell Lab of Ornithology. doi:10.2173/bna.682.
- ^ Angus, Wilson. "Identification and range of subspecies within the Canada and Cackling Goose Complex (Branta canadensis & B. hutchinsii)". http://www.oceanwanderers.com/CAGO.Subspecies.html.
- ^ "Chicago Area Is Home to Growing Numbers of Coyotes". Illinois Department of Natural Resources. http://dnr.state.il.us/orc/wildlife/virtual_news/releases/070104_coyotes.htm. Retrieved 2011-04-23.
- ^ a b [bna.birds.cornell.edu/bna/species/682/articles/behavior|Canada Goose behavior] (2011).
- ^ Johnsgard, Paul A. (2010) [1978]. Ducks, Geese, and Swans of the World (revised online ed.). Lincoln, Nebraska: University of Nebraska Press. p. 79. http://digitalcommons.unl.edu/biosciducksgeeseswans/5/.
- ^ Pennsylvania Department of Conservation and Natural Resources. Retrieved December 15, 2007, http://www.dcnr.state.pa.us/
- ^ Woodruff, Roger A.; Green, Jeffrey S. (1995). "Livestock Herding Dogs: A Unique Application for Wildlife Damage Management". Great Plains Wildlife Damage Control Workshop Proceedings (April 10–13, Ardmore, OK: Noble Foundation) 12: 43–45
- ^ Board of Park Commissioners (Seattle) Meeting Minutes July 12, 2001
- ^ Gregg MacDonald, Fairfax County Times (May 6, 2008). "Goose egg addling stirs concern in Reston". http://www.restontimes.com/news/2008/may/06/goose-egg-addling-stirs-concern-reston/. Retrieved 2009-06-10.
- ^ Division of Wildlife (Ohio) Goose Attacks
- ^ http://www.af.mil/news/airman/1297/bash.htm Air Force News article on Yukla 27
- ^ Barbara Barrett (2009-06-08). "DNA shows jet that landed in Hudson struck migrating geese". http://www.mcclatchydc.com/homepage/story/69645.html. Retrieved 2009-06-08.
- ^ Maynard, Micheline (15 January 2009). "Bird Hazard Is Persistent for Planes". The New York Times. http://www.nytimes.com/2009/01/16/nyregion/16strike.html?hp. Retrieved 23 February 2010.
- ^ T. M. John & J. C. George (1978). "Circulatory levels of thyroxine (T4) and triiodothyronine (T3) in the migratory Canada goose". Physiological Zoology 51 (4): 361–370. JSTOR 30160961.
- ^ Mėta M. Landys, John C. Wingfield & Marilyn Ramenofsky (2004). "Plasma corticosterone increases during migratory restlessness in the captive white-crowned sparrow Zonotrichia leucophrys gambelli" (PDF). Hormones and Behavior 46 (5): 574–581. doi:10.1016/j.yhbeh.2004.06.006. PMID 15555499. http://folk.uio.no/larsejo/tits/documents/Landys%20et%20al.%202004c.pdf.
Unreviewed
Names and Taxonomy
Taxonomy
Comments: Various authorities recognize 8-12 or more subspecies. Generally tundra populations comprise small birds, whereas southern birds are much larger, with intermediate-sized birds in intermediate localities. Within these size groupings, birds from the Pacific coast tend to be much darker than are eastern birds. However, reintroductions from mixed stocks have greatly muddied the traits of many southern populations. (DeBenedictus, Birding, Dec. 1991). Northern populations of small Canada Goose have been variously treated as three separate species: B. hutchinsii, B. minima, and B. leucopareia; as a single species, B. hutchinsii; or as one or more subspecies of B. canadensis (AOU 1983). B. hutchinsii is now recognized based on genetic studies, including recent work with mitochondrial DNA (AOU 2004). Mitochondrial DNA data indicate that Canada goose subspecies fall clearly into two sister groups, large-bodied and small-bodied, which share no mtDNA types (Van Wagner and Baker 1990, Quinn et al. 1991); in contrast, nuclear-gene-encoded protein evidence associates all the subspecies very closely (Van Wagner and Baker 1986, 1990).
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