Overview

Brief Summary

Flying fish, family Exocoetidae, are a diverse group of about 60+ fish in 7 genera (some classifications claim up to 9 genera). These mainly pelagic marine fish are widespread and abundant in the tropical and subtropical areas of the Pacific, Atlantic and Indian oceans. Flying fish primarily eat zooplankton, and are in turn an important food source for many marine predators including dolphinfishes, tunas, billfishes, cetaceans, as well as pelagic seabirds. Although they can’t actually fly, they are well known for their enlarged pectoral fins (“wings”), which allow them to make gliding leaps out of the water, a behavior believed to help them escape predation. Some species of flying fish have enlarged pelvic fins as well as enlarged pectoral fins, which allows them to fly further than two winged gliders (up to 400 meters), and have far greater maneuverability. A recent molecular phylogenetic analysis of the flying fish supports previous hypotheses that the evolution of four-winged gliding evolved once from a two-winged ancestor, perhaps in a three-step progression of gliding abilities (Lewallen et al 2011; Kutschera 2005). Flying fish have evolved a diversity of reproductive and life histories strategies: some lay their eggs in the open ocean, and have buoyant eggs that float on the ocean surface or non-buoyant eggs that have stringy filaments which get wound up in floating debris, others spend their lives in coastal areas, or return to coastal areas to breed; the different species also show a great diversity in how wide a range they occupy.

Flying fish fuel important commercial fisheries in Asia and are also commonly fished in other places, including the Caribbean. In Barbados, flying fish were threatened by pollution and overfishing, changing the occurrences of flying fish in the waters off of Barbados. This sparked a fishing controversy between Barbados, where flying fish is a traditional delicacy, and Trinidad and Tobago. As well as the fish meat, flying fish roe is commonly collected and is a common ingredient in sushi.

(Lewallen et al. 2011; Kutschera 2005; Potts et al. 2003; Wikipedia 2012)

  • Lewallen, E.A., R.L. Pitman, S.L. Kjartanson, N.R. Lovejoy, 2011. Molecular systematics of flyingfishes ( Teleostei : Exocoetidae ): evolution in the epipelagic zone. Fisheries Science Volume: 102, Issue: 1, Publisher: Blackwell Publishing Ltd, Pages: 161-174.
  • Kutschera, U. 2005. Predator-driven macroevolution in flyingfishes inferred from behavioural studies: historical controversies and a hypothesis. Annals of the History and Philosophy of Biology 10: 59–77.
  • Potts, A. C., Thomas, A. D., & Nichols, E. 2003. An economic and social assessment of the flying fish (pelagic) fishery of Tobago, Trinidad and Tobago. Proceedings of the Fifty Fourth Annual Gulf and Caribbean Fisheries Institute, 0-635.
  • Wikipedia, The Free Encyclopedia. 11 March, 2012. “Flying fish”. Retrieved March 15, 2012 from http://en.wikipedia.org/w/index.php?title=Flying_fish&oldid=481395284
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Ecology

Associations

Known predators

Exocoetidae (flying fishes) is prey of:
Gempylus serpens
Cephalopoda
Coryphaena

Based on studies in:
unknown (epipelagic zone, Tropical)

This list may not be complete but is based on published studies.
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Known prey organisms

Exocoetidae (flying fishes) preys on:
Euphausiidae
Copepoda
Hyperiidae

Based on studies in:
unknown (epipelagic zone, Tropical)

This list may not be complete but is based on published studies.
  • N. V. Parin, Ichthyofauna of the Epipelagic Zone (Israel Program for Scientific Translations, Jerusalem, 1970; U.S. Department of Commerce Clearinghouse for Federal Scientific and Technical Information, Springfield, VA 22151), from p. 154.
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Wikipedia

Flying fish

Exocoetidae, is a family of marine fish in the order Beloniformes of class Actinopterygii. Fish of this family are known as flying fish. There are about sixty-four species grouped in seven to nine genera. Flying fish can make powerful, self-propelled leaps out of water into air, where their long, wing-like fins enable them to glide for considerable distances above the water's surface. This uncommon ability is a natural defense mechanism to evade predators.

Contents

Etymology

The term "Exocoetidae" is not only the present scientific name for a genus of flying fish in this family, but also the general name in Latin for a flying fish. The suffix -idae, common for indicating a family, follows the root of the Latin word exocoetus, a transliteration of the Ancient Greek name ἐξώκοιτος. This means literally "sleeping outside", from ἔξω}"outside" and κοῖτος "bed", "resting place",[1], so named as flying fish were believed to leave the water to sleep on the shore.[2]

Flying fish have in turn given their name to:

Distribution and description

Flying fish live in all of the oceans, particularly in tropical and warm subtropical waters. Their most striking feature is their pectoral fins,[4] which are unusually large, and enable the fish to hide and escape from predators[5] by leaping out of the water and flying through air a few feet above the water's surface. Their flights are typically around 50 meters (160 ft).[6]

Flying fish taking off

To glide upward out of the water, a flying fish moves its tail up to 70 times per second.[7] It then spreads its pectoral fins and tilts them slightly upward to provide lift.[4] At the end of a glide, it folds its pectoral fins to reenter the sea, or drops its tail into the water to push against the water to lift itself for another glide, possibly changing direction.[4][7] The curved profile of the "wing" is comparable to the aerodynamic shape of a bird wing.[8] The fish is able to increase its time in the air by flying straight into or at an angle to the direction of updrafts created by a combination of air and ocean currents.[4][7]

Genus Exocoetus has one pair of fins and a streamlined body to optimize for speed, while Cypselurus has a flattened body and two pairs of fins which maximizes its time in the air. From 1900 to the 1930s, flying fish were studied as possible models used to develop airplanes.[7]

Exocoetidae feed mainly on plankton. Predators include dolphins, tuna, marlin, birds, squids and porpoises.[7]

Flight measurements

In May 2008, a Japanese television crew (NHK) filmed a flying fish (dubbed "Icarfish") off the coast of Yakushima Island, Japan. The creature spent 45 seconds in flight.[9] The previous record was 42 seconds.[9]

Flying fish can use updrafts at the leading edge of waves to cover distances of at least 400 m (1,300 ft).[6] They can travel at speeds of more than 70 kilometres per hour (43 mph).[7] Maximum altitude is 6 m (20 ft) above the surface of the sea.[8] Some accounts have them landing on ships' decks.[7][10]

Fishery and cuisine

Tobiko in varying colors. The bright orange roe is the natural coloration.

Flying fish are commercially fished in Japan, Vietnam and China by the method of gillnetting, and in Indonesia and India by dipnetting.[7] In Japanese cuisine, the fish is often preserved by drying. The roe of Cheilopogon agoo, or Japanese flying fish, is used to make some types of sushi, and is known as tobiko. It is also a staple in the diet of the Tao people of Orchid Island, Taiwan. Flying fish is part of Barbados' national dish,[11] known as cou-cou and flying fish.[12][13]

In the Solomon Islands, they are caught while flying, using nets held from outrigger canoes. They are attracted to the light of torches. Fishing is done only when there is no moonlight.

Importance

Barbados

Historically, Barbados was nicknamed "the land of the flying fish", where today it is the official national fish.[14] Once abundant, it migrated between the warm, coral-filled Atlantic Ocean surrounding the island of Barbados and the plankton-rich outflows of the Orinoco River in Venezuela.

Just after the completion of the Bridgetown Harbor / Deep Water Harbor in Bridgetown, Barbados saw an increase of ship visits, linking the island to the world. The overall health of the coral reefs surrounding Barbados suffered due to ship-based pollution. Additionally, Barbadian overfishing pushed them closer to the Orinoco river delta, no longer returning to Barbados in large numbers. Today, the flying fish only migrate as far north as Tobago, around 120 nautical miles (220 km; 140 mi) southwest of Barbados. Despite the change, flying fish remain a coveted delicacy.

Many aspects of Barbadian culture center around the flying fish: it is depicted on coins, as sculptures in fountains, in artwork, and as part of the official logo of the Barbados Tourism Authority. Additionally, the Barbadian coat of arms features a pelican and dolphin fish on either side of the shield, but the dolphin resembles a flying fish. Furthermore, actual artistic renditions and holograms of the flying fish are also present within the Barbadian passport.

Maritime disputes

In recent times, flying fish have also been gaining in popularity in other islands, fueling several maritime disputes. In 2006, the council of the United Nations Convention on the Law of the Sea[15] fixed the maritime boundaries between Barbados and Trinidad and Tobago over the flying fish dispute, which gradually raised tensions between the neighbours.[16] The ruling stated both countries must preserve stocks for the future. Barbadian fishers still follow the flying fish southward. Flying fish remain an important part of Barbados' main national dish.

See also

References

  1. ^ Harper, Douglas. "exocet". Online Etymology Dictionary. http://www.etymonline.com/index.php?term=exocet. 
  2. ^ Pliny's Natural History, vol. IX, 19)
  3. ^ Guillot, Jean; Estival, Bernard (1988). L’extraordinaire aventure de l’Exocet. Les éditions de la Cité. ISBN 2-85186-039-9.  The missile's name was given by M. Guillot, then technical director at Nord Aviation, after the French name for flying fish.
  4. ^ a b c d Fish, F. E. (1990). "Wing design and scaling of flying fish with regard to flight performance" (PDF). Journal of Zoology 221 (3): 391–403. doi:10.1111/j.1469-7998.1990.tb04009.x. http://darwin.wcupa.edu/~biology/fish/pubs/pdf/1990JZWingdesign.pdf. 
  5. ^ Buller, D. J. (1998). "Etiological theories of function: a geographical survey" (PDF). Biology and Philosophy 13 (4): 505–527. doi:10.1023/A:1006559512367. http://www.niu.edu/phil/~buller/research/etf.pdf. 
  6. ^ a b Ross Piper (2007), Extraordinary Animals: An Encyclopedia of Curious and Unusual Animals, Greenwood Press.
  7. ^ a b c d e f g h Kutschera, U. (2005). "Predator-driven macroevolution in flyingfishes inferred from behavioural studies: historical controversies and a hypothesis" (PDF). Annals of the History and Philosophy of Biology 10: 59–77. http://www.evolutionsbiologen.de/flyingfishes.pdf. 
  8. ^ a b Fish, F. (1991). "On a fin and a prayer" (PDF). Scholars 3 (1): 4–7. http://darwin.wcupa.edu/~biology/fish/pubs/pdf/1991SOnafin.pdf. 
  9. ^ a b "Fast flying fish glides by ferry". BBC News. May 20, 2008. http://news.bbc.co.uk/1/hi/sci/tech/7410421.stm. Retrieved May 20, 2008. 
  10. ^ Joseph Banks (1997). The Endeavour Journal of Sir Joseph Banks 1768-1771. University of Sydney Library. http://setis.library.usyd.edu.au/ozlit/pdf/p00021.pdf. Retrieved July 16, 2009. 
  11. ^ Flying fish of Barbados :: Gowealthy.com
  12. ^ "Editorial: Our Fisheries — rights and duties". Trinidad and Tobago Express. June 25, 2006. http://www.trinidadexpress.com/index.pl/article?id=160973250. 
  13. ^ "Tribunal reaches decision in the marine dispute between Trinidad & Tobago and Barbados". Trinidad and Tobago Express. http://www.trinidadexpress.com/index.pl/article?id=150421244. 
  14. ^ "Outstanding Barbadians & Cultural Symbols - The Flying Fish". Central Bank of Barbados. http://centralbank.org.bb/WEBCBB.nsf/webpage/96C400BBEED7A84B042572FC0018B1CD?OpenDocument#Flyingfish. 
  15. ^ "Barbados/Trinidad and Tobago". Permanent Court of Arbitration. April 11, 2006. http://www.pca-cpa.org/showpage.asp?pag_id=1152. 
  16. ^ "Claims of Caribbean piracy as national symbol takes flight". The Sydney Morning Herald. December 18, 2004. http://www.smh.com.au/news/World/Claims-of-Caribbean-piracy-as-national-symbol-takes-flight/2004/12/17/1102787277205.html. 
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