Two subfamilies are recognized (Nielsen 1978, Davis 1990, 1998): Eriocottinae, which are mostly Old World from southern Palearctic of Spain and North Africa (Eriocottis, 10 sp., Deuterotinea, 5 sp.) to Australia (Eucryptogona, 1 sp.), with a single genus in South America (Crepidochares, 5 sp.), and Compsocteninae containing a single genus ranging from southern Africa to Taiwan (Compsoctena, ca. 50 sp.).
Nearctic, Palearctic, Oriental, Ethiopian, Neotropical, Australian
Adult Abdomen Morphology
Female genitalia description:
Segments 8 and 9 + 10 forming a slender, elongate, telescoping oviscapt supported by typically 2 pairs of elongate apophyses and 2 (rarely 3) pairs of pseudapophyses (Davis 1990a); anterior dorsal pseudapophyses located entirely within A8 and usually fused for most of their length. Ostium situated at caudal margin of membranous S8. Ductus bursae elongate, membranous. Corpus bursae membranous; signum a single, irregular sclerotization, or absent.
Female pregenital sexual scales:
Female accessory glands:
Female oviduct opening:
Female bursa ostium opening:
on venter 8
Female anterior apophyses originating:
from venter 8
Male pregenital sexual scales:
Male genitalia description:
Uncus moderately bilobed, fused to hoodlike tegumen. Vinculum usually short and V-shaped, rarely Y-shaped (with an elongate saccus). Gnathos slender, U- to V-shaped or reduced and widely separated. Subscaphium moderately sclerotized. Transtilla present. Juxta usually indistinct, slender and weakly sclerotized in Eriocottis. Valvae typically with a prominent spinose lobe (clasper) from medioventral margin, or with a more basal, variably developed process from sacculus (in Crepidochares). Aedoeagus a simple, slender cylinder without cornuti.
Sternum 5 gland:
Adult abdomen description:
Paired sternal apodemes of A2 slender, continuing half way into sternum as sternal rods.
Adult Thorax Morphology
Adult thorax description:
Metafurca with apophyses free.
Number of tibial spurs foreleg:
Number of tibial spurs midleg:
Number of tibial spurs hindleg:
Epiphysis relatively smooth (without pecten) in Compsoctena.
Forewing length from mid-dorsum to apex (mm):
from 4.5 to 23
All veins arising separate from cell; Rs4 terminating either on costa or termen; forewing with both accessory and intercalary cells preserved; a retinaculum consisting of a broad fold arising ventrally from costal margin and partially extending over base of Sc: I A + 2 A with basal fork.
Forewing cell veins:
Forewing basal loop:
present, with frenulum
Wings moderately broad, forewing index 0.23-0.4; microtrichia sparsely scattered over all wing surfaces; Female of Deuterotinea brachypterous (Zagulayev 1988);
Hindwings about as broad as forewings; hindwing index 0.32-0.4; male frenulum a single stout seta; female with 2-4 smaller frenular setae.
Adult Head Morphology
porrect, upcurved, large
Number of labial palp segments:
Number of maxillary palp segments:
from 2 to 5
Proboscis moderate to elongate in subfamily Eriocottinae; minute in Compsoctinae.
Number of flagellomere scale rows:
General antennae description:
Antenna 0.4-0.7 the length of forewing; pecten present; flagellum usually filiform, bipectinate in ♂ Compsoctena, with one row of slender bidentate scales per segment, either restricted to dorsal surface or completely encircling segment, more concentrated in Compsoctena.
Adult head description:
Vestiture rough consisting of mostly long piliform scales with simple, acute apices. Ocelli present or absent. Eye small to moderately large; interocular index 0.6-1.1; cornea with sparse scattering of interfacetal microsetae.
Usually 2 (rarely 3) pairs of pseudapophyses within female oviscapt. Anterior dorsal pseudapophyses located entirely within A8 and usually fused for most of their length
Molecular Biology and Genetics
Statistics of barcoding coverage
Specimens with Sequences:106
Specimens with Barcodes:103
Species With Barcodes:7
Eriocottidae or Old World spiny-winged moths is a family of insects in the Lepidoptera order whose position relative to other members of the superfamily Tineoidea is currently unknown. There are between eight and seventeen (Natural History Museum Lepidoptera genus database) genera currently placed within it and two subfamilies, Compsocteninae and Eriocottinae.
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