Overview

Brief Summary

Diversity

Diversity description:

Two subfamilies are recognized (Nielsen 1978, Davis 1990, 1998): Eriocottinae, which are mostly Old World from southern Palearctic of Spain and North Africa (Eriocottis, 10 sp., Deuterotinea, 5 sp.) to Australia (Eucryptogona, 1 sp.), with a single genus in South America (Crepidochares, 5 sp.), and Compsocteninae containing a single genus ranging from southern Africa to Taiwan (Compsoctena, ca. 50 sp.).

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Distribution

Geographical Distribution

Geographic Range:

Nearctic, Palearctic, Oriental, Ethiopian, Neotropical, Australian

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Physical Description

Morphology

Adult Abdomen Morphology

Reproductive system:

Ditrysian

Oviscapt (ovipositor):

non-piercing

Female genitalia description:

Segments 8 and 9 + 10 forming a slender, elongate, telescoping oviscapt supported by typically 2 pairs of elongate apophyses and 2 (rarely 3) pairs of pseudapophyses (Davis 1990a); anterior dorsal pseudapophyses located entirely within A8 and usually fused for most of their length. Ostium situated at caudal margin of membranous S8. Ductus bursae elongate, membranous. Corpus bursae membranous; signum a single, irregular sclerotization, or absent.

Female corethrogyne:

absent

Female pregenital sexual scales:

absent

Female accessory glands:

one pair

Female oviduct opening:

below anus

Female bursa ostium opening:

on venter 8

Female anterior apophyses originating:

from venter 8

Male coremata:

absent

Male pregenital sexual scales:

absent

Male genitalia description:

Uncus moderately bilobed, fused to hoodlike tegumen. Vinculum usually short and V-shaped, rarely Y-shaped (with an elongate saccus). Gnathos slender, U- to V-shaped or reduced and widely separated. Subscaphium moderately sclerotized. Transtilla present. Juxta usually indistinct, slender and weakly sclerotized in Eriocottis. Valvae typically with a prominent spinose lobe (clasper) from medioventral margin, or with a more basal, variably developed process from sacculus (in Crepidochares). Aedoeagus a simple, slender cylinder without cornuti.

Sternum 5:

without fenestra

Sternum 5 gland:

absent

Adult abdomen description:

Paired sternal apodemes of A2 slender, continuing half way into sternum as sternal rods.

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Adult Thorax Morphology

Scale tufts:

present

Epiphysis:

present

Adult thorax description:

Metafurca with apophyses free.

Forelegs:

normal

Number of tibial spurs foreleg:

from 0

Number of tibial spurs midleg:

from 2

Number of tibial spurs hindleg:

from 4

Leg description:

Epiphysis relatively smooth (without pecten) in Compsoctena.

Forewing length from mid-dorsum to apex (mm):

from 4.5 to 23

Wing venation??description:

All veins arising separate from cell; Rs4 terminating either on costa or termen; forewing with both accessory and intercalary cells preserved; a retinaculum consisting of a broad fold arising ventrally from costal margin and partially extending over base of Sc: I A + 2 A with basal fork.

Wing venation:

heteroneurous

Forewing cell veins:

forked

Forewing basal loop:

present

Wing coupling:

present, with frenulum

Wing scales:

hollow

Forewing description:

Wings moderately broad, forewing index 0.23-0.4; microtrichia sparsely scattered over all wing surfaces; Female of Deuterotinea brachypterous (Zagulayev 1988);

Hindwing description:

Hindwings about as broad as forewings; hindwing index 0.32-0.4; male frenulum a single stout seta; female with 2-4 smaller frenular setae.

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Tympanum Morphology

Counter-tympanum:

absent

Abdomen tympanum:

absent

Thorax tympanum:

absent

Palp tympanum:

absent

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Adult Head Morphology

Ocelli:

present, absent

Eyes:

smooth

Labial palpus:

porrect, upcurved, large

Number of labial palp segments:

from 3

Maxillary palpus:

present

Number of maxillary palp segments:

from 2 to 5

Proboscis:

present, reduced

Proboscis texture:

naked

Proboscis description:

Proboscis moderate to elongate in subfamily Eriocottinae; minute in Compsoctinae.

Mandibles:

absent, reduced

Female antennae:

filiform

Male antennae:

bipectinate, filiform

Number of flagellomere scale rows:

from 1

General antennae description:

Antenna 0.4-0.7 the length of forewing; pecten present; flagellum usually filiform, bipectinate in ♂ Compsoctena, with one row of slender bidentate scales per segment, either restricted to dorsal surface or completely encircling segment, more concentrated in Compsoctena.

Adult head description:

Vestiture rough consisting of mostly long piliform scales with simple, acute apices. Ocelli present or absent. Eye small to moderately large; interocular index 0.6-1.1; cornea with sparse scattering of interfacetal microsetae.

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Diagnostic Description

Synapomorphies

Apomorphies:

Usually 2 (rarely 3) pairs of pseudapophyses within female oviscapt.  Anterior dorsal pseudapophyses located entirely within A8 and usually fused for most of their length

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Molecular Biology and Genetics

Molecular Biology

Statistics of barcoding coverage

Barcode of Life Data Systems (BOLD) Stats
Specimen Records: 97
Specimens with Sequences: 95
Specimens with Barcodes: 92
Species: 8
Species With Barcodes: 7
Public Records: 11
Public Species: 4
Public BINs: 8
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Barcode data

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Wikipedia

Eriocottidae

Eriocottidae or Old World spiny-winged moths[1] is a family of insects in the Lepidoptera order whose position relative to other members of the superfamily Tineoidea is currently unknown. There are between eight[2] and seventeen (Natural History Museum Lepidoptera genus database) genera currently placed within it and two subfamilies, Compsocteninae and Eriocottinae.

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