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Distribution: Spain (dunes of Coto Doñana, south of Andalusia, more than 400 km south of the Sistema Central range (Magraner, 1986); this was often refuted (Peréz-Mellado, 1997, 1998; Barbadillo et al., 1999); mountains to the Sistema Central and along the Atlantic lowlands (Sá-Sousa 1999, 2000). berlengensis: Berlengas islands off the western coast of Portugal (Vicente, 1985);
Type locality: Berlenga Island, Portugal.
Type locality: Laguna de San Marco, La Alberca, Prov. Salamanca, Spain.