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 Life habit: lichenized, in some cases lichenicolous in young developmental stages; Thallus: thickly crustose, rimose-areolate, sometimes placodioid, or subsquamulose; areoles: flat or convex to almost bullate; upper surface: smooth, dull or subnitid, whitish-gray pruinose, or naked and brown; upper cortex: paraplectenchymatous, composed of small roundish-angular cells; medulla: white, subparaplectenchymatous; lower and lateral cortices: where present as in upper cortex; photobiont: primary one a chlorococcoid alga, secondary photobiont absent; algal layer: continuous; the algal cells: ± in columns; lower surface: attached with ± constricted bases or stipes, or rhizine-like structures; rhizohyphae: lacking; Ascomata: perithecial, laminal, immersed, subglobose; exciple: colorless or brown to black; involucrellum: lacking (or a very reduced apical lid); interascal filaments: absent; ostiolar filaments: present; asci: clavate, wall not amyloid, 8-spored; ascospores: biseriate, ellipsoid or oblong-ovoid, hyaline, simple or 1-, rarely 3-septate, non-halonate, smooth; Conidiomata: pycnidial, laminal, immersed, Dermatocarpon-type; conidia: subcylindrical to bacilliform; Secondary metabolites: none detected; Geography: world-wide, particularly in temperate, semi-arid regions; Substrate: calciferous or non-calcareous rock. 

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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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