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DescriptionThallus: determinate, effigurate to rimose areolate, orbicular or not, often confluent, up to 6 cm or more across,; areoles: irregularly shaped, 0.5-1 mm across, mostly fertile, mostly densely contiguous and distinctly radiating, margin: with lobes 1.5-2.3 mm. long and as wide; upper surface: bright yellow, shiny, rough and becoming rugose; upper cortex: 40-70 µm thick, mostly yellow; algal layer: 40-90 µm thick, continuous or becoming irregularly interrupted by hyphal bundles, with an uneven surface; medulla: intricate to anticlinal prosoplectenchyma with granules, thickness varying, without a lower surface, continuous with attaching hyphae; Apothecia: less than 1 mm in diam.; disc: usually yellow, plane to very convex; margin: never completely excluded; true exciple: parathecium: often forming parathecial crown, up to 50 µm thick,; epihymenium: dark greenish yellow, 20 µm thick; hymenium: mostly dirty or pale yellow, 65-100 µm tall; paraphyses: conglutinate, with many septae, 1.7-2 µm thick at base; tips: not swollen; subhymenium: hyaline, 50-80(-170) µm thick, with mainly anticlinal hyphae; hypothecium: hyaline, 30-50 µm thick, the hyphae parallel or intricate; asci: clavate, 50-70 x 16-24 µm, 100+-spored; ascospores: hyaline, rather narrowly ellipsoid, simple, 4-5 x 1.7-2 µm; Pycnidia: deeply immersed with pale ostioles; conidia: 2-2.5 x 0.7-0.8 µm; Spot tests: thallus UV+ orange, K-, C-, KC-. P-; Secondary metabolites: containing rhizocarpic, acaranoic and acarenoic or lichesterinic acid .; Substrate and ecology: on perpendicular or overhanging hard acid rock surfaces, usually middle-level montane to alpine; World distribution: Europe and North America; Sonoran distribution: Arizona, southern California, Baja California, Baja California Sur, Chihuahua, and Sonora.; Notes: Variation in the species is wider than most descriptions suggest, leading to some confusion. Sometimes all or part of the thallus is smooth, sometimes the margin becomes much reduced and the disc can become very convex and appear biatorine but is always less than 1 mm wide.