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 Thallus: strongly squamulose, areolate; prothallus: indistinct; areoles: dark brown to black on the underside contiguous to dispersed, flat, irregularly ear-shaped in outline, 0.5-1.5 mm in diam.; surface: light brown with gray margins, esorediate; medulla: white, I-; Apothecia: black, sessile, slightly constricted at base, up to 2.5 mm in diam.; disc: flat to moderately convex, dull, epruinose or indistinctly pruinose; margin: black, persistent, dull to shiny; exciple: marginally with a thin, 15-20 µm wide, black to dark brown rim and a hyaline to dull beige (by crystalline immersion) interior part, peripherally 30-80 µm wide, K-, C-, P-; epihymenium: dark brown with a weak olive tinge, c. 10 µm thick; hymenium: hyaline, (65-)73-84(-95) µm tall, I+ orange; paraphyses: simple, occasionally branched and anastomosing, c. 2 µm in diam., with slightly swollen apical cells of 3-4 µm in diam.; subhymenium: unpigmented, (20-)25-35(-45) µm thick; hypothecium: dark brown to almost black; asci: clavate, Lecidea-type, 55-75 x 12-16 µm, 8-spored; ascospores: hyaline, simple, ellipsoid, (9.5-)13-15 (-19) x (4.5-)5-6.5(-8) µm, length-widthindex 2.1-2.3; Pycnidia: immersed, graphidoid; conidia: cylindrical, 7.5-11 x 1-1.3 µm [studied in 2 specimens]; Spot tests: cortex and medulla K-, C-, KC-, P-; Secondary metabolites: perlatolic acid.; Substrate and ecology: on exposed acidic rocks in the lower montane zone; World distribution: only known in its type collection; Sonoran distribution: not recorded; to be expected southwards in California.; Notes: Lecidea brodoana is related to Lecidea fuscoatra. It has a rather pronounced squamulous thallial areolation (a character, which, although less pronounced, sometimes is also found in the very variable L. fuscoatra) and possesses perlatolic acid instead of gyrophoric acid. It is named in honor of the great lichenologist Irwin M. Brodo. 

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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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