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 Squamules: up to 6 mm wide, elongate, adnate and dispersed to adjacent or imbricate, weakly concave to weakly convex; upper surface: castaneous brown, shiny, epruinose, smooth or sparingly fissured; margin: concolorous with upper side, often slightly down-turned, entire, crenulate or lobed; upper cortex: up to 140 µm thick, composed of thick-walled hyphae with angular lumina, containing crystals of lichen substances but no calcium oxalate; medulla: not containing crystals; lower cortex: of mainly periclinally oriented hyphae, not containing calcium oxalate; lower surface: medium brown, K-; Apothecia: up to 1.5 mm diam., laminal, immarginate or marginate when young and later immarginate, dark brown to black, epruinose; ascospores: ellipsoid, 12-14 x 5-7 µm; Pycnidia: laminal, immersed; conidia: bacilliform, 5-7 x 1 µm; Spot tests: upper cortex K-, C+ red, KC+ red, P-; medulla K-, C-, KC-, P-; Secondary metabolites: bourgeanic and gyrophoric acids, and trace of lecanoric acid.; Substrate and ecology: on soil and in crevices of rock, in open habitats in the lowlands, up to 980 m alt. in California, at 1250 m on Isla Guadalupe; World distribution: west coast of North America north to Washington; Sonoran distribution: fairly common in coastal California and Baja California.; Notes: The C+ red and KC+ red reactions in the upper cortex are often difficult to observe. The present delimitation conforms with chemical strain I of Timdal (1986); strain II is here treated as P. hyporubescens. 


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© Lichen Unlimited: Arizona State University, Tempe.

Source: Lichen Flora of the Greater Sonoran Desert Region

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