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 Thallus: squamulose; squamules: c. 2-4 mm wide and 0.35 mm thick, fully appressed, roundish or slightly lobed, ± dispersed or a few adjacent; upper surface: brown, dull; upper cortex: c. 30-60 µm thick, epinecral layer almost lacking or up to 20 µm thick; medulla: white, usually with numerous globose cells (c. 9-14 µm in diam.); algal layer: c. 70-100 µm high; lower cortex: absent or at least lacking distinctive development; lower surface: brownish; rhizohyphae: hyaline, rather thin (3.5-4.5 µm); Perithecia: broadly pyriform to subglobose, up to 0.45 mm wide, with colorless wall; asci: cylindrical, 70-80 x 13-17 µm, 8-spored; ascospores: uniseriate, subglobose or broadly ellipsoid, 10-13 (-15) x (7.5-) 8-9 (-10) µm, rather thick-walled; Pycnidia: not seen; Spot tests: all negative; Secondary metabolites: none detected.; Substrate and ecology: soil, in open places, mostly coastal; World distribution: endemic to SW North America; Sonoran distribution: on coastal sites of southern California and Baja California.; Notes: The species very much resembles P. squamulosum, but differs in its almost spherical and thick-walled spores and in having thinner rhizohyphae. 

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Source: Lichen Flora of the Greater Sonoran Desert Region

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