Physical Description

Type Information

Isotype for Lysiloma kellermanii Britton & Rose
Catalog Number: US 2442699
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Original publication and alleged type specimen examined
Preparation: Pressed specimen
Collector(s): W. A. Kellerman
Year Collected: 1907
Locality: El Rancho, Jalapa, Guatemala, Central America
Elevation (m): 1000 to 1000
  • Isotype: Britton, N. L. & Rose, J. N. 1928. N. Amer. Fl. 23: 81.
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Type collection for Lysiloma calderonii Britton & Rose
Catalog Number: US 1151978
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Card file verified by examination of alleged type specimen
Preparation: Pressed specimen
Collector(s): S. Calderón
Year Collected: 1922
Locality: Cerro de La Olla, near Chalchuapa., Santa Ana, El Salvador, Central America
  • Type collection: Britton, N. L. & Rose, J. N. 1925. N. Amer. Fl. 28: 82.
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Type collection for Lysiloma australis Britton & Rose
Catalog Number: US 1080750
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Original publication and alleged type specimen examined
Preparation: Pressed specimen
Collector(s): C. Conzatti
Year Collected: 1907
Locality: El Parian., Nochixtlan, Oaxaca, Mexico, North America
Elevation (m): 1000 to 1000
  • Type collection: Britton, N. L. & Rose, J. N. 1928. N. Amer. Fl. 23: 83.
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Type collection for Lysiloma chiapensis Britton & Rose
Catalog Number: US 1209403
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Original publication and alleged type specimen examined
Preparation: Pressed specimen
Collector(s): C. A. Purpus
Year Collected: 1925
Locality: Banks of creek W of Monserrate., Chiapas, Mexico, North America
  • Type collection: Britton, N. L. & Rose, J. N. 1928. N. Amer. Fl. 23: 83.
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Type collection for Lysiloma salvadorensis Britton & Rose
Catalog Number: US 1137014
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Original publication and alleged type specimen examined
Preparation: Pressed specimen
Collector(s): P. C. Standley
Year Collected: 1922
Locality: Vicinity of San Vicente., San Vicente, El Salvador, Central America
Elevation (m): 350 to 500
  • Type collection: Britton, N. L. & Rose, J. N. 1928. N. Amer. Fl. 23: 83.
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Ecology

Habitat

Sierra de la Laguna Dry Forests Habitat

This taxon is found in the Sierra de la Laguna dry forests ecoregion, which was once an isolated island, containing a large number of endemic species. After sufficient mountain uplift and the joining the Baja Peninsula mainland, this ecoregion underwent significant speciation, and is thus today high in species diversity; this portion of the peninsula contains the majority of the species found in the southern part of the Baja Peninsula. The region is shaped by a vast complex of granitic mountains, running southward from the Gulf of California to the Pacific. These mountains are dissected by valleys and canyons, and surrounded by vast plateaus.

The forest is transitional both with the pine oak forests at higher elevations, and with the xeric scrub at lower portions. The dry forest of Sierra de la Laguna is characterized by abundance of low trees and scrubs, and poor vertical stratification. The dominant tree species in the subtropical forest are Mauto (Lysiloma divaricatum), Palo Blanco (L. candida), Elephant Tree (Bursera microphylla) and Palo Zorrillo (Hesperalbizia occidentalis). Herbaceous elements are poorly developed, but their representatives are Caribe (Cnidoscolus angustidens), Spiny Aster (Chloracantha spinosa var. strictospinosa), Solanum spp., and cacti such as Biznaga (Ferocactus spp).

A number of reptilian taxa are found in the ecoregion, including: the endemic Baja California Rat Snake (Bogertophis rosaliae); Hunsaker's Spiny Lizard (Sceloporus hunsakeri); Belding's Orange-throated Whiptail (Cnemidophorus hyperythrus); Spiny Chuckwalla (Sauromalus hispidus NT); San Lucan Leaf-tailed Gecko (Phyllodactylus unctus NT); Baja California Night Snake (Hypsiglena slevini), a Mexican endemic rangeing from Bahía San Juanico, in the east-central Baja California Peninsula, southward continuously Cabo San Lucas (as well as on the island of Santa Margarita and on Cerralvo and Danzante islands in the Gulf of California; and Hunsaker's Spiny Lizard (Sceloporus hunsakeri), endemic to the Cape Region of Baja California Sur and the Gulf of California islands of Espiritu Santo, Gallo, Ballena and Partida Sur.

There are a number of mammalian species occurring in the Sierra de la Laguna dry forests. Among the mammals found here are: Eva's Desert Mouse (Peromyscus eva), endemic to Baja California Sur;  Mexican Funnel-eared Bat (Natalus stramineus); the near-endemic Peninsular Bat (Myotis peninsularis EN), chiefly found in Baja California Sur; Dalquest's Pocket Mouse (Chaetodipus dalquesti VU), known only from the Cape Region of Baja California Sur.

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San Lucan Xeric Scrub Habitat

This taxon is found in the San Lucan xeric scrub, an ecoregion situated at the southern-most part of the Baja Peninsula of Mexico; this diverse landscape of mountains, valleys, and plateaus is covered with a variety of species of xeric vegetation. This neotropical ecoregion is classifed within the Deserts and Xeric Scrublands biome. Plants and animals of this region evolved independently before the Baja Peninsula, a previous island during the Miocene, joined the mainland. An arid climate supports a number fauna and species, about ten percent which of which are endemic.

The ecoregion took shape in the Miocene as an isolated landform prior to joining the peninsula, and thus can be considered an biogeographical island of vegetation. This arid landscape is composed of a vast, rugged complex of granitic mountains, valleys, canyons, and plateaus. The ecoregion occupies the plateaus between the coast and the lower limits of the dry forests, which begin around 250 meters. Precipitation is about 400 millimetres annually.

Some elements of dry forest habitat are present in this ecoregion, but xeric elements are dominant and include Chain-link Cholla (Opuntia cholla); Elephant Tree (Bursera microphylla), at the southern limit of its range here and extending north to the Waterman Mountains in the USA; Mauto (Lysiloma divaricata); Organ Pipe Cactus (Stenocereus thuberi), Mala Mujer (Cnidoscolus angustidens), Yucca spp., and Barrel Cacti (Ferocactus spp). Herbaceous elements in the ecoregion include Plantago linearis, Bouteloua hirsuta, and Commelina coelestis.

The San Lucan xeric scrub harbours 31 of 48 of the reptile species for the Cape Region. Almost a third of the wider regional recorded species of collembola arthropods and spiders (30 of 138 species, respectively) occur in this ecoregion. In general, over ten percent of animal and plant species found here are endemic.

Within the San Lucan xeric scrub ecoregion, reptilian taxa include: the endemic Island Burrowing Sand Snake (Chilomeniscus punctatissimus); the endemic Isla Cerralvo Snake (Chilomeniscus savagei), restricted solely to Cerralvo Island; the Cape Arboreal Spiny Lizard (Sceloporus licki), a near-endemic restricted to the southern portion of the Baja Peninsula; the near-endemic Spiny Chuckwalla (Sauromalus hispidus NT), found only on Angel de la Guarda Island, Granito, Mejía, Pond, San Lorenzo Norte, San Lorenzo Sur, and other islands in Bahía de los Ángeles, including Cabeza de Caballo, La Ventana, Piojo, Flecha, Mitlàn, and Smith, Gulf of California; the near-endemic San Lucan Leaf-toed Gecko (Phyllodactylus unctus NT), found only on southern Baja Peninsula and some islands within the Gulf of California: Gallo, Partida Sur, Espiritu Santo, Ballena, Gallina and Cerralvo. There are only a small number of anuran species present in the ecoregion: Red-spotted Toad (Anaxyrus punctatus); and Pacific Chorus Frog (Pseudacris regilla).

The Espiritu Santo Island Antelope Squirrel (Ammospermophilus insularis) is endemic to the San Lucan xeric scrub ecoregion and is found only on the island of Espiritu Santo in the Gulf of California. Among threatened mammals occurring in the ecoregion are: the near-endemic Dalquest's Pocket Mouse (Chaetodipus dalquesti VU), known from the Cape Region of the Baja California Peninsula.

Threatened mammals in the ecoregion include: the near-endemic Peninsular Myotis (Peninsular Myotis EN), found only on southern Baja Peninsula; Fish-eating Bat (Myotis vivesi VU), a near-endemic occurring chiefly on the near-shore islands off of the southern Baja Peninsula and mainland Sonora; Mexican Long-tongued Bat (Choeronycteris mexicana NT); and the Lesser Long-nosed Bat (Leptonycteris yerbabuenae VU).

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Southern Pacific Dry Forests Habitat

This taxon is found in the Southern Pacific dry forests ecoregion, which is situated along the southeastern versant of the Sierra Madre del Sur Mountains including the Pacific Ocean coastal plain. These forests are a key locus of endemism for butterflies, and has the greatest diversity of scorpions and spiders in the entirety of Mexico. This ecoregion is classified in the Tropical and Subtropical Dry Broadleaf Forests biome. The Southern Pacific dry forests exhibit a moderate to high faunal species richness; for example, there are a total of 744 vertebrate taxa recorded in the ecoregion, with a particularly large number of endemic reptiles.

The ecoregion elevation ranges from sea level to 1400 metres. The climate is tropical and dry, with precipitation levels of 800 millimetres (mm) per annum. There is an extended arid season, which factor drives the prevalence of deciduous vegetation. The forests grow chiefly on shallow, well-drained soils derived from limestone. Closer to the base of the Sierra Madre del Sur Mountains, the soils are more rocky, and are derived from igneous rocks.

The dominant plant species include Mauto (Lysiloma divaricatum), Bursera excelsa and Fragrant Bursera (B. fagaroides), which are typically found in association with Pochote (Ceiba aesculifolia), Comocladia engleriana, and Trichilia americana. In the Mexican state of Michoacán, the macro plant species more generally in evidence are Ficus insipida, F. pertusa, Breadnut (Brosimum alicastrum), Licania arborea, Sideroxylon capiri and Elephant Ear (Enterolobium cyclocarpum).

There are a number of anuran species present in the ecoregion, including: Blunt-toed Chirping Frog (Eleutherodactylus modestus VU); Cloud Forest Stream Frog (Ptychohyla euthysanota NT), found from southeast Oaxaca to Guatemala and eastern El Salvador; Matuda's Spikethumb Frog  (Plectrohyla matudai VU). A special status caecilian found in the ecoregion is the Mexican Caecilian (Dermophis mexicanus VU), a fossorial species that can attain lengths up to sixty centimetres. A special status salamander found in the ecoregion is the Sierra Juarez Salamander (Pseudoeurycea juarezi CR), a near-endemic known only between Cerro Pelón and Vista Hermosa in the Sierra de Juarez, north-central Oaxaca. The White-lipped Peeping Frog (Eleutherodactylus albolabris CR), a near-endemic known chiefly from Agua del Obispo, central Guerrero.

The Southern Pacific dry forests contain numerous reptilian taxa, including the following endemics: Bocourt's Anole (Norops baccatus); Taylor's Anole (Norops taylori), known only to  Puerto Marquez area, in northern Acapulco, Guerrero; Simmons' Anole (Anolis simmonsi), restricted to the vicinity of Pinotepa Nacional, Oaxaca; Stegneger's Blackcollar Spiny Lizard (Sceloporus stejnegeri), restricted to the Pacific versant in the state of Guerrero, Mexico; Red Earth Snake (Geophis russatus), found in a very narrow range outside of Putla, Oaxaca; Sierra Mije Earth Snake (Geophis anocularis), known only from around Totontepec on the Atlantic versant of the Sierra Mixe, Oaxaca; Ramirez`s Hooknose Snake (Ficimia ramirezi), restricted to the Pacific versant of the Isthmus of Tehuantepec, Niltepec, Oaxaca; Halberg's Cloud Forest Snake (Cryophis hallbergi), found only in northern Oaxaca, at Sierra de Juarez and Sierra Mazateca; Isthmian Earth Snake (Geophis isthmicus), known only from the vicinity of Tehuantepec, Mexico; the endemic Macdougall's Spiny Lizard (Sceloporus macdougalli).

Characteristic mammalian fauna include the endemic Oaxacan Pocket Gopher (Orthogeomys cuniculus), restricted to several sites on the Isthmus of Tehuantepec, Oaxaca. Other mammals seen in the ecoregion include the: Lesser Long-nosed Bat (Leptonycteris yerbabuenae VU), Tropical Hare (Lepus flavigularis EN), restricted to Salina Cruz, Oaxaca to the extreme west of  Chiapas; Greater Bulldog Bat (Noctilio leporinus), Coati (Nasua narica), Buller’s Pocket Gopher (Pappogeomys bulleri), Javelina (Tayassu tajacu), and Mexican Long-tongued Bat (Choeronycteris mexicana NT).

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Molecular Biology and Genetics

Molecular Biology

Statistics of barcoding coverage: Lysiloma divaricatum

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
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