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Cheilanthes — Details

Lip Ferns learn more about names for this taxon

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Overview

Comprehensive Description

Description

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Rhizome erect or shortly creeping, with linear scales. Stipe dark reddish-brown to black, glabrous and glossy or pilose. Fronds 2-5-pinnatifid, uniform or rarely dimorphic, the pinnae and ultimate segments never articulated, glabrous, pubescent, pilose or tomentose, but never with coloured powdery undumentum on the underside. Sori small, discrete, borne on vein endings near the margins, or in soral lines, protected by a continuous or interrupted indusium or by the recurved lobes of the ultimate segments.

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Distribution

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Pellaea marginata (Kunth) Baker:
Ecuador (South America)

Sodiro, L. 1893. Crypt. Vasc. Quit. 1–656. Typis Universitatis, Quito. http://www.tropicos.org/Reference/1000624

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Cheilanthes Sw.:
Argentina (South America)
Belize (Mesoamerica)
Bolivia (South America)
Brazil (South America)
Chile (South America)
Colombia (South America)
Costa Rica (Mesoamerica)
Ecuador (South America)
El Salvador (Mesoamerica)
Guatemala (Mesoamerica)
Honduras (Mesoamerica)
Nicaragua (Mesoamerica)
Panama (Mesoamerica)
Peru (South America)
Paraguay (South America)
Uruguay (South America)
United States (North America)
Venezuela (South America)
Mexico (Mesoamerica)
Caribbean (Caribbean)

Forzza, R. C. & et al. 2010. 2010 Lista de espécies Flora do Brasil. http://floradobrasil.jbrj.gov.br/2010/. http://www.tropicos.org/Reference/100002289
Molina Rosito, A. 1975. Enumeración de las plantas de Honduras. Ceiba 19(1): 1–118. http://www.tropicos.org/Reference/866
Luteyn, J. L. 1999. Páramos, a checklist of plant diversity, geographical distribution, and botanical literature. Mem. New York Bot. Gard. 84: viii–xv, 1–278. http://www.tropicos.org/Reference/1024098
Davidse, G., M. Sousa Sánchez & S. Knapp. (eds.) 1995. Psilotaceae a Salviniaceae. Fl. Mesoamer. 1: i–xxi, 1–470. http://www.tropicos.org/Reference/47044

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Allosorus ciliatus var. minor C. Presl:
Mexico (Mesoamerica)

SPECIMEN BASED RECORD. Published protolog data. http://www.tropicos.org/Reference/9990002

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Cheilanthes nudiuscula (R. Br.) T. Moore:
China (Asia)

Flora of China Editorial Committee. 1988-2013. Fl. China Unpaginated. Science Press & Missouri Botanical Garden Press, Beijing & St. Louis. http://www.tropicos.org/Reference/42480

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Onychium angustifolium Ching:
China (Asia)

Flora of China Editorial Committee. 1988-2013. Fl. China Unpaginated. Science Press & Missouri Botanical Garden Press, Beijing & St. Louis. http://www.tropicos.org/Reference/42480

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Cheilanthes venusta var. aurata Fée:
Mexico (Mesoamerica)

SPECIMEN BASED RECORD. Published protolog data. http://www.tropicos.org/Reference/9990002

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Allosorus angustifolius var. minus Liebm.:
Mexico (Mesoamerica)

SPECIMEN BASED RECORD. Published protolog data. http://www.tropicos.org/Reference/9990002

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Cheilanthes pyramidalis Fée:
Costa Rica (Mesoamerica)
El Salvador (Mesoamerica)
Guatemala (Mesoamerica)
Honduras (Mesoamerica)
Mexico (Mesoamerica)
Nicaragua (Mesoamerica)

SPECIMEN BASED RECORD. Published protolog data. http://www.tropicos.org/Reference/9990002
Molina Rosito, A. 1975. Enumeración de las plantas de Honduras. Ceiba 19(1): 1–118. http://www.tropicos.org/Reference/866
Pfeiffer-Berendsohn, B. 1989. Pteridophyta in: Listado Basico de la Flora Salvadorensis. Cuscatlania 1: 1–28. http://www.tropicos.org/Reference/11333
Mickel, J. T. & A. R. Smith. 2004. The Pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1–1054. http://www.tropicos.org/Reference/1025841

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Pellaea angustifolia var. elongata Rovirosa:
Mexico (Mesoamerica)

SPECIMEN BASED RECORD. Published protolog data. http://www.tropicos.org/Reference/9990002

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Cheilanthes hirsuta Link:
Colombia (South America)
Costa Rica (Mesoamerica)
El Salvador (Mesoamerica)
Guatemala (Mesoamerica)
Honduras (Mesoamerica)
Mexico (Mesoamerica)
Nicaragua (Mesoamerica)
Panama (Mesoamerica)

SPECIMEN BASED RECORD. Published protolog data. http://www.tropicos.org/Reference/9990002
Pfeiffer-Berendsohn, B. 1989. Pteridophyta in: Listado Basico de la Flora Salvadorensis. Cuscatlania 1: 1–28. http://www.tropicos.org/Reference/11333
Mickel, J. T. & J. M. Beitel. 1988. Pteridophyte Flora of Oaxaca, Mexico. Mem. New York Bot. Gard. 46: 1–568. http://www.tropicos.org/Reference/9402
Smith, A. R. 1981. Pteridophytes. 2: 1–370. In D. E. Breedlove Fl. Chiapas. California Academy of Sciences, San Francisco. http://www.tropicos.org/Reference/6014
Correa A., M. D., C. Galdames & M. N. S. Stapf. 2004. Cat. Pl. Vasc. Panamá 1–599. Smithsonian Tropical Research Institute, Panama. http://www.tropicos.org/Reference/1031911
Breedlove, D. E. 1986. Flora de Chiapas. Listados Floríst. México 4: i–v, 1–246. http://www.tropicos.org/Reference/513
Luteyn, J. L. 1999. Páramos, a checklist of plant diversity, geographical distribution, and botanical literature. Mem. New York Bot. Gard. 84: viii–xv, 1–278. http://www.tropicos.org/Reference/1024098
Davidse, G., M. Sousa Sánchez & S. Knapp. (eds.) 1995. Psilotaceae a Salviniaceae. Fl. Mesoamer. 1: i–xxi, 1–470. http://www.tropicos.org/Reference/47044

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Cheilanthes cuneata Kaulf. ex Link:
Costa Rica (Mesoamerica)
Guatemala (Mesoamerica)
Mexico (Mesoamerica)
Venezuela (South America)

Mickel, J. T. & A. R. Smith. 2004. The Pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1–1054. http://www.tropicos.org/Reference/1025841
Mickel, J. T. & J. M. Beitel. 1988. Pteridophyte Flora of Oaxaca, Mexico. Mem. New York Bot. Gard. 46: 1–568. http://www.tropicos.org/Reference/9402
Smith, A. R. 1981. Pteridophytes. 2: 1–370. In D. E. Breedlove Fl. Chiapas. California Academy of Sciences, San Francisco. http://www.tropicos.org/Reference/6014
Hokche, O., P. E. Berry & O. Huber. 2008. Nuev. Cat. Fl. Vas. Venezuela 1–860. Fundación Instituto Botánico de Venezuela, Caracas, Venezuela. http://www.tropicos.org/Reference/1033110
Breedlove, D. E. 1986. Flora de Chiapas. Listados Floríst. México 4: i–v, 1–246. http://www.tropicos.org/Reference/513
Davidse, G., M. Sousa Sánchez & S. Knapp. (eds.) 1995. Psilotaceae a Salviniaceae. Fl. Mesoamer. 1: i–xxi, 1–470. http://www.tropicos.org/Reference/47044

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Cheilanthes angustifolia Kunth:
Colombia (South America)
Costa Rica (Mesoamerica)
El Salvador (Mesoamerica)
Guatemala (Mesoamerica)
Honduras (Mesoamerica)
Mexico (Mesoamerica)
Nicaragua (Mesoamerica)
Panama (Mesoamerica)
Venezuela (South America)

SPECIMEN BASED RECORD. Published protolog data. http://www.tropicos.org/Reference/9990002
Molina Rosito, A. 1975. Enumeración de las plantas de Honduras. Ceiba 19(1): 1–118. http://www.tropicos.org/Reference/866
Pfeiffer-Berendsohn, B. 1989. Pteridophyta in: Listado Basico de la Flora Salvadorensis. Cuscatlania 1: 1–28. http://www.tropicos.org/Reference/11333
Mickel, J. T. & A. R. Smith. 2004. The Pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1–1054. http://www.tropicos.org/Reference/1025841
Mickel, J. T. & J. M. Beitel. 1988. Pteridophyte Flora of Oaxaca, Mexico. Mem. New York Bot. Gard. 46: 1–568. http://www.tropicos.org/Reference/9402
Fournier, E. 1870. Sur deux fougères nouvelles du Nicaragua. Bull. Soc. Bot. France 17: 236–237. http://www.tropicos.org/Reference/1006007
Smith, A. R. 1981. Pteridophytes. 2: 1–370. In D. E. Breedlove Fl. Chiapas. California Academy of Sciences, San Francisco. http://www.tropicos.org/Reference/6014
Correa A., M. D., C. Galdames & M. N. S. Stapf. 2004. Cat. Pl. Vasc. Panamá 1–599. Smithsonian Tropical Research Institute, Panama. http://www.tropicos.org/Reference/1031911
Breedlove, D. E. 1986. Flora de Chiapas. Listados Floríst. México 4: i–v, 1–246. http://www.tropicos.org/Reference/513
Davidse, G., M. Sousa Sánchez & S. Knapp. (eds.) 1995. Psilotaceae a Salviniaceae. Fl. Mesoamer. 1: i–xxi, 1–470. http://www.tropicos.org/Reference/47044
Stolze, R. G. 1981. Ferns and fern allies of Guatemala. Part II. Polypodiaceae. Fieldiana, Bot., n.s. 6: 1–522. http://www.tropicos.org/Reference/6644

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Cheilanthes marginata Kunth:
Argentina (South America)
Bolivia (South America)
Colombia (South America)
Costa Rica (Mesoamerica)
Ecuador (South America)
El Salvador (Mesoamerica)
Guatemala (Mesoamerica)
Honduras (Mesoamerica)
Mexico (Mesoamerica)
Panama (Mesoamerica)
Peru (South America)
Venezuela (South America)

Molina Rosito, A. 1975. Enumeración de las plantas de Honduras. Ceiba 19(1): 1–118. http://www.tropicos.org/Reference/866
Foster, R. C. 1958. A catalogue of the ferns and flowering plants of Bolivia. Contr. Gray Herb. 184: 1–223. http://www.tropicos.org/Reference/1313
González, J. A., G. J. Scrocchi & E. O. Lavilla. 1999. Relevamiento de la biodiversidad de la reserva nacional de flora y fauna Tariquía (Tarija, Bolivia). Ser. Conserv. Natur. 14: 7–134. http://www.tropicos.org/Reference/1012020
Adolfo M., H. 1966. Nóm. Pl. Recol. Valle Cochabamba 2: 17–86. Colegio La Salle, Cochabamba. http://www.tropicos.org/Reference/1018799
Pfeiffer-Berendsohn, B. 1989. Pteridophyta in: Listado Basico de la Flora Salvadorensis. Cuscatlania 1: 1–28. http://www.tropicos.org/Reference/11333
Mickel, J. T. & A. R. Smith. 2004. The Pteridophytes of Mexico. Mem. New York Bot. Gard. 88: 1–1054. http://www.tropicos.org/Reference/1025841
Mickel, J. T. & J. M. Beitel. 1988. Pteridophyte Flora of Oaxaca, Mexico. Mem. New York Bot. Gard. 46: 1–568. http://www.tropicos.org/Reference/9402
Kessler, M., T. Krömer & I. Jiménez. 2000. Inventario de grupos selectos de plantas en el valle de Masicurí (Santa Cruz--Bolivia). Revista Boliviana Ecol. Cons. Amb. 8: 3–15. http://www.tropicos.org/Reference/1016408
Correa A., M. D., C. Galdames & M. N. S. Stapf. 2004. Cat. Pl. Vasc. Panamá 1–599. Smithsonian Tropical Research Institute, Panama. http://www.tropicos.org/Reference/1031911
Hokche, O., P. E. Berry & O. Huber. 2008. Nuev. Cat. Fl. Vas. Venezuela 1–860. Fundación Instituto Botánico de Venezuela, Caracas, Venezuela. http://www.tropicos.org/Reference/1033110
Jørgensen, P. M. & S. León-Yánez. (eds.) 1999. Catalogue of the vascular plants of Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75: i–viii, 1–1181. http://www.tropicos.org/Reference/42250
Breedlove, D. E. 1986. Flora de Chiapas. Listados Floríst. México 4: i–v, 1–246. http://www.tropicos.org/Reference/513
Luteyn, J. L. 1999. Páramos, a checklist of plant diversity, geographical distribution, and botanical literature. Mem. New York Bot. Gard. 84: viii–xv, 1–278. http://www.tropicos.org/Reference/1024098
Davidse, G., M. Sousa Sánchez & S. Knapp. (eds.) 1995. Psilotaceae a Salviniaceae. Fl. Mesoamer. 1: i–xxi, 1–470. http://www.tropicos.org/Reference/47044

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Molecular Biology and Genetics

Molecular Biology

Statistics of barcoding coverage: Cheilanthes pyramidalis

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Barcode of Life Data Systems (BOLDS) Stats
Public Records: 14
Species: 14
Species With Barcodes: 1

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Statistics of barcoding coverage: Cheilanthes angustifolia

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Barcode of Life Data Systems (BOLDS) Stats
Public Records: 12
Species: 12
Species With Barcodes: 1

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Statistics of barcoding coverage: Cheilanthes cuneata

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Barcode of Life Data Systems (BOLDS) Stats
Public Records: 4
Species: 4
Species With Barcodes: 1

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Statistics of barcoding coverage: Cheilanthes marginata

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Barcode of Life Data Systems (BOLDS) Stats
Public Records: 18
Species: 18
Species With Barcodes: 1

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Wikipedia

Cheilanthes

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Cheilanthes (lip ferns)^[2] is a genus of about 180 species of rock-dwelling ferns with a cosmopolitan distribution in warm, dry, rocky regions, often growing in small crevices high up on cliffs. Most are small, sturdy and evergreen. The leaves, often densely covered in trichomes, spring directly from the rootstocks. Many of them are desert ferns, curling up during dry times and reviving with the coming of moisture. At the ends of veins sporangia, or spore-bearing structures, are protected by leaf margins, which curl over them.
This genus is now known to be highly paraphyletic, comprising at least four generically separate groups. The type species, C. micropteris, is most closely allied to the genera Aleuritopteris and Sinopteris (Schuettpelz et al.).
The genus name is derived from the Greek words χεῖλος (cheilos), meaning "lip," and ἄνθος (anthos), meaning "flower."^[3]
Selected species

Cheilanthes acrostica (Balb.) Tod.
Cheilanthes adiantoides T.C.Chambers & P.A.Farrant
Cheilanthes aemula Maxon – Rival lip fern
Cheilanthes alabamensis (Buckley) Kunze – Alabama lip fern
Cheilanthes arizonica (Maxon) Mickel – Arizona lip fern
Cheilanthes austrotenuifolia H.M.Quirk & T.C.Chambers
Cheilanthes bonariensis (Willd.) Proctor – Golden lip fern
Cheilanthes brownii (Kuhn) Domin
Cheilanthes catanensis (Cosent.) H.P.Fuchs
Cheilanthes caudata R.Br.
Cheilanthes cavernicola D.L.Jones 1988 (Australia)^[4]
Cheilanthes clevelandii D.C.Eaton – Cleveland's lip fern
Cheilanthes contigua Baker
Cheilanthes cooperae D.C.Eaton – Cooper's lip fern
Cheilanthes covillei Maxon – Coville's lip fern
Cheilanthes distans (R.Br.) Mett. – Bristly cloak fern
Cheilanthes eatonii Baker – Eaton's lip fern
Cheilanthes feei T.Moore – Slender lip fern
Cheilanthes fendleri Hook. – Fendler's lip fern
Cheilanthes fragillima F.Muell.
Cheilanthes glauca (Cav.) Mett.
Cheilanthes gracillima D.C.Eaton – Lace lip fern
Cheilanthes guanchica Bolle
Cheilanthes hirsuta (Poir.) Mett.
Cheilanthes hispanica Mett.
Cheilanthes horridula Maxon – Rough lip fern
Cheilanthes humilis (G.Forst.) P.S.Green
Cheilanthes intertexta (Maxon) Maxon – Coastal lip fern
Cheilanthes intramarginalis (Kaulf. ex Link) Hook.
Cheilanthes kaulfussii Kunze – Kaulfuss' lip fern
Cheilanthes lanosa (Michx.) D.C.Eaton
Cheilanthes lasiophylla Pic.Serm.
Cheilanthes lendigera (Cav.) Sw. – Nit-bearing lip fern
Cheilanthes leucopoda Link – Whitefoot lip fern
Cheilanthes lindheimeri Hook. – Lindheimer lip fern
Cheilanthes maderensis Lowe
Cheilanthes microphylla (Sw.) Sw. – Southern lip fern
Cheilanthes micropteris Cheilanthes newberryi (D.C.Eaton) Domin – Newberry's lip fern
Cheilanthes nitida (R.Br.) P.S.Green
Cheilanthes nudiuscula (R.Br.) T.Moore
Cheilanthes parryi (D.C.Eaton) Domin – Parry's lip fern
Cheilanthes paucijuga Baker
Cheilanthes peninsularis
Cheilanthes peninsularis var. insularis
Cheilanthes persica (Bory) Mett. ex Kuhn
Cheilanthes pinnatifida D.L.Jones
Cheilanthes praetermissa D.L.Jones
Cheilanthes prenticei Luerss.
Cheilanthes pringlei Davenport – Pringle's lip fern
Cheilanthes pseudovellea (H.M.Quirk & T.C.Chambers) D.L.Jones

Cheilanthes pteroides Cheilanthes pulchella Bory ex Willd.
Cheilanthes pumilio (R.Br.) F.Muell.
Cheilanthes sciadioides Domin
Cheilanthes sieberi Kunze – Poison rock fern
Cheilanthes tenuifolia (Burm.f.) Sw.
Cheilanthes tinaei Tod.
Cheilanthes tomentosa Link – Woolly lip fern
Cheilanthes vellea (Aiton) F.Muell.
Cheilanthes villosa Davenport ex Maxon – Villous lip fern
Cheilanthes viscida Davenport – Viscid lip fern
Cheilanthes wootonii Maxon – Beaded lip fern
Cheilanthes wrightii Hook. – Wright's lip fern
Cheilanthes yavapensis T.Reeves ex Windham – Graceful lip fern^[2]

Formerly placed here

Astrolepis cochisensis (Goodd.) D.M.Benham & Windham (as C. cochisensis (Goodd.) Mickel and C. sinuata var. cochisensis (Goodd.) Munz)
Notholaena candida (M.Martens & Galeotti) Hook. (as C. candida M.Martens & Galeotti)
Pteris lydgatei (Baker) Christ (as C. lydgatei Baker)^[5]

References

Wikimedia Commons has media related to: Cheilanthes

Wikispecies has information related to: Cheilanthes

^ ^a ^b "Genus: Cheilanthes Sw.". Germplasm Resources Information Network. United States Department of Agriculture. 2007-10-05. http://www.ars-grin.gov/cgi-bin/npgs/html/genus.pl?13099. Retrieved 2011-06-07.
^ ^a ^b "Cheilanthes". Integrated Taxonomic Information System. http://www.itis.gov/servlet/SingleRpt/SingleRpt?search_topic=TSN&search_value=17432. Retrieved 2011-10-23.
^ Moran, Robbin Craig (2004). A Natural History of Ferns. Timber Press. p. 94. ISBN 978-0-88192-667-5. http://books.google.com/books?id=zy1hcSGaaDAC&.
^ C. cavernicola International Plant Names Index (IPNI) 13 Jan 2012
^ "GRIN Species Records of Cheilanthes". Germplasm Resources Information Network. United States Department of Agriculture. http://www.ars-grin.gov/cgi-bin/npgs/html/splist.pl?13099. Retrieved 2011-06-07.

"Cheilanthes Sw.". Australian Plant Name Index (APNI), IBIS database. Centre for Plant Biodiversity Research, Australian Government. http://www.anbg.gov.au/cgi-bin/apni?taxon_id=7697.
Flora Europaea: Cheilanthes
Flora of North America: Cheilanthes
Cheilanthes glauca pictures from Chilebosque.
"lip fern." Encyclopædia Britannica Online. 30 Nov 2007.

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