Canada

Origin: Unknown/Undetermined

Regularity: Regularly occurring

Currently: Unknown/Undetermined

Confidence: Confident

United States

Origin: Unknown/Undetermined

Regularity: Regularly occurring

Currently: Unknown/Undetermined

Confidence: Confident

Cryptogramma acrostichoides has often been treated as a variety or subspecies of the strictly European Cryptogramma crispa (Linneaus) R. Brown, which has a chromosome number of 2 n = 120.

Stems decumbent to erect, much branched from base, stout, 10--20 mm diam. (including hardened, persistent leaf bases); scales bicolored, dense, broadly lanceolate to linear, to 6 × 2 mm. Leaves densely tufted, green over winter, persistent after withering; sterile leaves spreading, 3--17 cm; fertile leaves erect, 5--25 cm; hairs small, appressed, cylindric, scattered along grooves of petiole and along costae and costules of adaxial blade surface. Petiole green to straw-colored, dark brown on proximal 1/8 or less, 1--2 mm wide, firm and strawlike, not collapsed; scales bicolored or ± concolored, becoming sparse distally. Blade deltate to ovate-lanceolate, all 2--3-pinnate, somewhat leathery, opaque; hydathodes sunken below leaf surface. Segments of sterile leaves oblong to ovate-lanceolate, bases cuneate, distal 2/3--1/2 of segments crenate to dentate, often somewhat more deeply incised every 2d tooth; segments of fertile leaves horizontal to ascending, strongly differentiated from those of sterile leaves, linear, 3--12 × 1--2 mm; margins of fertile segments revolute, covering sporangia. Sporangia in sori that coalesce at maturity. 2 n = 60.

Cryptogramma crispa (Linnaeus) R. Brown ex Hooker subsp. acrostichoides (R. Brown) Hultén; C. crispa var. acrostichoides (R. Brown) C. B. Clarke

Comments: Crevices, ledges, and talus slops and in pockets of organic soil in the Precambrian region of northern North America (Cody, 1989).

New growth produced in spring, spores maturing in summer, sterile leaves green over winter, senescing 2d spring. Noncalcareous cliff crevices, rock outcrops, and talus, often in relatively dry habitats, typically montane but occurring in lowland to alpine habitats; 0--3700 m; Alta., B.C., Man., N.W.T., Ont., Sask., Yukon; Alaska, Ariz., Calif., Colo., Idaho, Mich., Mont., Minn., Nev., N.Mex., Oreg., Utah, Wash., Wyo.; reported from Mexico in Baja California; Asia.

Canada

Rounded National Status Rank: NNR - Unranked

United States

Rounded National Status Rank: NNR - Unranked

Rounded Global Status Rank: G5 - Secure

Reasons: Common through much of Canada and southern Alaska, extending down into California, New Mexico, Minnesota and Michigan.

Comments: Scoggan's (1978) Cryptogramma crispa ssp. acrostichoides var. acrostichoides is a synonym of C. acrostichoides since Kartesz (1994) treats the other variety, var. sitchensis, as a distinct species.