Cheilanthes cooperae — Details

Cheilanthes cooperae D.C. Eaton:
United States (North America)

Anonymous. 1986. List-Based Rec., Soil Conserv. Serv., U.S.D.A. Database of the U.S.D.A., Beltsville. http://www.tropicos.org/Reference/1103
Lellinger, D. B. 1985. A Field Manual of the Ferns and Fern Allies of the United States and Canada. 389 pp. http://www.tropicos.org/Reference/1316
Munz, P. A. & D. D. Keck. 1959. Cal. Fl. 1–1681. University of California Press, Berkeley. http://www.tropicos.org/Reference/1717
Munz, P. A. 1974. Fl. S. Calif. 1–1086. University of California Press, Berkeley. http://www.tropicos.org/Reference/1719

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National Distribution

United States

Origin: Unknown/Undetermined

Regularity: Regularly occurring

Currently: Unknown/Undetermined

Confidence: Confident

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Physical Description

Morphology

Comments

Although scattered throughout much of California, Cheilanthes cooperae is apparently rare and quite localized. It appears to be most closely related to C . viscida , from which it differs in lacking glandular pubescence. The ranges of the two species do not overlap, and they seem amply distinct.

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Description

Stems compact to short-creeping, usually 4--8 mm diam.; scales brown at base and darker toward tip, linear-subulate, straight to slightly contorted, loosely appressed, persistent. Leaves clustered, 5--30 cm; vernation circinate. Petiole dark brown, flattened or slightly grooved distally on adaxial surface. Blade linear-oblong to lanceolate-ovate, 3-pinnate at base, 1.5--5 cm wide; rachis flattened or slightly grooved adaxially, lacking scales, with monomorphic pubescence. Pinnae not articulate, dark color of stalk continuing into pinna base, basal pair slightly smaller than adjacent pair, ± equilateral, appearing hirsute adaxially. Costae green adaxially for most of length; abaxial scales absent. Ultimate segments linear-oblong to ovate, not beadlike, the largest 3--5 mm, abaxially and adaxially hirsute with long, strongly flattened hairs. False indusia marginal, weakly differentiated, 0.05--0.25 mm wide. Sori usually discontinuous, concentrated on apical and lateral lobes. Sporangia containing 64 spores. 2 n = 60.

Flora of North America @ eFloras.org

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Type Information

Type collection for Cheilanthes cooperae D.C. Eaton
Collection: Smithsonian Institution, National Museum of Natural History, Department of Botany
Verification Degree: Verified from the card file of type specimens
Preparation: Pressed specimen
Collector(s): E. Cooper
Locality: California, United States, North America

Type collection: Eaton, D. C. 1875. Bull. Torrey Bot. Club. 6: 33.

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Ecology

Habitat

Habitat & Distribution

Sporulating late spring--summer. Calcareous cliffs and ledges; usually on limestone; 100--700 m; Calif.

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Conservation

Conservation Status

NatureServe Conservation Status

Rounded Global Status Rank: G3 - Vulnerable

Source: NatureServe

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National NatureServe Conservation Status

United States

Rounded National Status Rank: NNR - Unranked

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Wikipedia

Cheilanthes cooperae

Cheilanthes cooperae is a species of lip fern known by the common name Mrs. Cooper's lip fern, or simply Cooper's lip fern. It is endemic to California, where it is widespread but not very common. It can be found growing in crevices in rocky habitat, generally on limestone.

Description

The leaves are up to about 30 centimeters long and a few wide, and are made up of segments subdivided into pairs of many-lobed smaller segments. They are pale green in color with glandular hairs. The smallest segments are oval in shape and have sori with tan-colored sporangia on their undersides.

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