Morphology

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Description

Stems short-creeping, usually 4--8 mm diam.; scales uniformly black or with narrow brown margins, linear-subulate, straight to slightly contorted, strongly appressed, persistent. Leaves clustered, 8--35 cm; vernation circinate. Petiole dark brown, flattened or slightly grooved distally on adaxial surface. Blade elongate-pentagonal, 3-pinnate-pinnatifid to 4-pinnate at base, 3--10 cm wide; rachis grooved adaxially, lacking scales, with monomorphic pubescence. Pinnae not articulate, dark color of stalk continuing into pinna base, basal pair larger than adjacent pair, strongly inequilateral, proximal basiscopic pinnules greatly enlarged, appearing glandular-pubescent adaxially. Costae green or straw-colored adaxially for most of length; abaxial scales absent. Ultimate segments linear-oblong, not especially beadlike, largest 3--8 mm, abaxially and adaxially glandular pubescent with short, sticky, capitate glands. False indusia marginal, weakly differentiated, 0.05--0.25 mm wide. Sori usually discontinuous, concentrated on apical and lateral lobes. Sporangia containing 32 spores.

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© Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA

Source: Missouri Botanical Garden

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