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Description

 Slender, low-climbing or trailing, perennial vine 1–3 m or more, minutely antrorsely appressed-puberulent throughout with unicellular, curved trichomes, 0.1–0.2 mm long, 0.02–0.03 mm wide. Flowering stems 1.3–2.3 mm in diameter, terete or somewhat compressed. Stipules 1.0–4.9 mm long, 0.3–0.7 mm wide, narrowly ovate, acute to slightly attenuate, longitudinally striate-nerved; petioles (0.1-)0.5–3.0 cm long, inserted 1.4–7.3(-8.9) mm from the basal margins of the peltate blades, with 2, round or elliptic, opposite to subopposite, sessile, discoid nectaries with flat rims, 1.1–1.9 mm wide (on the widest axis), 0.3–1.0 mm high, borne in the distal third of the petiole (0.63–0.87 of the distance from the base toward the apex of the petiole). Laminas 1.1–5.8 cm long, 3.4–13.7 cm wide, coriaceous, commonly variegated along primary veins and major secondary veins, conspicuously peltate, ratio of leaf width to central vein length measured from point of petiole insertion 2.0–5.9, depressed obovate to transversely elliptic (widely divaricately bilobed), lateral lobes 2.6–7.3(-8.1) cm long, 0.8–4.9 cm wide, elliptic, obtuse or acute to slightly attenuate, central lobe commonly obsolete or present as an obtuse to retuse tip, central vein 0.9–4.4(-5.4) cm long (measured from point of petiole insertion), angle between the lateral lobes 53–162°, ratio of lateral lobe to central vein length 1.3–3.0(-4.8), margins entire, hyaline, primary veins 3, diverging and branching above base, laminar nectaries present, 6–17, submarginal, associated with the minor veins of the abaxial surface, 0.8–1.3 mm in diameter, widely elliptic to circular, sessile; tendril 0.4–0.8 mm wide, present at flowering node, absent in inflorescence. Flowers borne in leaf axils or inflorescences; inflorescences 4.5–22.3 cm long, associated reduced laminas 2.1–5.3 mm long, 1.3–2.5 mm wide. Pedicels 1.3–3.1(-9.9) mm long, 0.4–1.1 mm wide, (1-)2 per node; bract(s) absent; spur(s) absent. Flowers 18.8–31.9 mm in diameter with stipe 12.3–19.0(-23.3) mm long, 0.4–0.8 mm wide; hypanthium 4.8–8.1 mm in diameter; sepals 6.5–12.3 mm long, 3.1–6.3 mm wide, ovate-triangular, acute, abaxially and adaxially greenish yellow; coronal filaments in 7 series, the outer 40–50, 6.3–10.4 mm long, 0.1–0.3 mm wide, linear, spreading flat, reflexed above middle and the tips often slightly incurved, very dark reddish purple with yellow at tips, ratio of outer coronal row to sepal length 0.73–1.38, the second 35–50, 2.3–4.8(-5.1) mm long, 0.1–0.2 mm wide, linear, spreading flat, very dark reddish purple with yellow tips, ratio of second coronal row to outer coronal row length 0.23–0.60, the third 40–50, 0.8–3.0 mm long, 0.06–0.13 mm wide, linear, spreading flat, very dark reddish purple with yellow tips, ratio of third coronal row to second coronal row length 0.20–0.65, the fourth through seventh ca. 100 per series, 0.6–1.3 mm long, 0.1–0.2 mm wide, linear, capitate, erect, very dark reddish purple, ratio of coronal rows 4–7 to third coronal row length 0.30–0.72(-0.91); operculum 0.3–0.7 mm long, denticulate, very dark reddish purple, nectary absent; limen absent, limen floor 4.7–7.1 mm in diameter, very dark reddish purple; androgynophore 2.7–4.1 mm long, 0.7–1.3 mm wide; free portions of the staminal filaments 2.3–3.6 mm long, 0.4–0.7 mm wide, linear, very dark reddish purple; anthers 1.6–3.1 mm long, 0.7–1.7 mm wide, extrorse at anthesis with their axes maintained parallel to the filament; styles 4.1–6.3 mm long including stigmas, 0.3–0.5 mm wide, greenish yellow with very dark reddish purple tinge; stigmas 0.7–1.6 mm in diameter; ovary 1.3–3.7 mm long, 1.4–2.7 mm wide, widely ellipsoid to globose, greenish yellow. Berry 14.4–26.0 mm long, 12.5–19.00 mm in diameter, widely ellipsoid to ovoid, greenish yellow with white spots, becoming soft at the base at maturity. Seeds ca. 10, 5.0–6.1 mm long, 2.1–2.7 mm wide, 1.3–1.9 mm thick, widely elliptic in outline, acute at both ends, reticulate-foveate with each face marked with 12–24 foveae. Germination type epigeal.

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© Kristen Porter-Utley

Source: PhytoKeys

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