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Slender, climbing, perennial vine 2–6 m long or more, sparsely pubescent with unicellular curved trichomes on petiole, leaf, stem, sepal, and stipule, 0.20–0.64 mm long, 0.02–0.03 mm wide, also minutely antrorsely appressed-puberulent throughout with unicellular, curved trichomes, 0.03–0.12 mm long, 0.02–0.03 mm wide. Flowering stems 1.0–2.4 mm in diameter, terete to somewhat compressed, with the base woody and cork-covered. Stipules 2.5–6.0 mm long, 0.4–1.3 mm wide, narrowly ovate-triangular, acute; petioles 1.2–5.7 cm long, with 2 (rarely 3), opposite to subopposite, sessile, discoid nectaries with flat rims, 1.0–2.1 mm wide (on the widest axis), 0.3–1.3 mm high, borne in the proximal half of the petiole (0.34–0.54 of the distance from the base toward the apex of the petiole). Laminas 2.6–8.5 cm long, 6.5–23.5 cm wide, coriaceous, peltate (the distance from leaf base to point of petiole insertion 3.0–18.9 mm), transversely elliptic (widely divaricately bilobed) or sometimes 3-lobed, lateral lobes 3.7–12.9 cm long, 1.9–7.5 cm wide, elliptic, acute to attenuate, central lobe elliptic to obovate or present merely as a widely acute to obtuse tip (rarely retuse), central vein 1.8–7.0 cm long (measured from point of petiole insertion to the leaf apex), angle between the lateral lobes 132–188°, ratio of lateral lobe to central vein length 1.33–2.77, margins entire, hyaline, primary veins 3, diverging and branching above base, laminar nectaries present, 4–13, submarginal, associated with the minor veins of the abaxial surface, with 2–4 nectaries proximal to the lateral leaf veins, 0.5–1.4 mm in diameter, circular to widely elliptic, sessile; juvenile leaves bilobed and variegated, the variegation seen in some clones at maturity; tendril 0.3–1.0 mm wide, present at flowering node, absent in inflorescence. Flowers borne in leaf axils or inflorescences; inflorescences 2.0–18.5(-25.1) cm long, associated reduced laminas 2.0–4.3 mm long, 1.5–3.1 mm wide. Pedicels 1.9–15.8 mm long, 0.4–0.9 mm wide, 2 per node; bract(s) absent; spur(s) absent. Flowers 18.4–33.4 mm in diameter with stipe 3.1–8.6(-9.4) mm long, 0.5–1.3 mm wide; hypanthium 5.4–8.2 mm in diameter; sepals 6.5–13.3 mm long, 2.9–6.3 mm wide, ovate-triangular, acute to rounded, abaxially and adaxially greenish yellow; coronal filaments in 2 series, the outer 40–51, 5.5–8.4 mm long, 0.3–0.7(-0.8)mm wide, linear, more or less erect, very dark reddish purple (5RP 3/2) on proximal third, greenish yellow (5GY 8/4) on middle third, yellow on distal third (5Y 8/10), ratio of outer coronal row to sepal length 0.59–0.94, the inner 27–40, 2.3–3.8 mm long, 0.2–0.5(-0.6) mm wide, linear, capitate, erect, greenish yellow with a flush of very dark reddish purple at base, ratio of inner coronal row to outer coronal row length 0.35–0.52; operculum 1.2–2.0 mm long, plicate, greenish yellow with a flush of very dark reddish purple at base, the margin white with narrow minutely fimbrillate teeth; nectary 0.1–0.5(-0.6) mm high, 0.5–1.2(-2.9) mm wide; limen not recurved but inclined toward the operculum, 0.1–0.5(-0.7) mm high, 0.1–0.4(-0.5) mm wide, very dark red (5R 2.5/2), limen floor 2.5–5.1 mm in diameter, very dark red; androgynophore (2.1-)3.5–6.0 mm long, 0.4–1.3(-1.5) mm wide, the distal half dark red then greenish yellow with dark red spots and streaks; free portions of the staminal filaments 1.5–2.8 mm long, 0.5–0.9 mm wide, linear, greenish yellow; anthers 1.9–2.9 mm long, 0.6–1.5(-1.9) mm wide; styles 1.8–4.9(-5.4) mm long including stigmas, 0.2–0.5 mm wide, greenish yellow; stigmas 0.8–1.7 mm in diameter; ovary 1.6–2.7 mm long, 1.1–2.1 mm wide, widely ellipsoid to globose, greenish yellow. Berry (12.4-)18–23.1 mm long, (13.4-)18.4–33.0 mm in diameter, globose, very dark purple. Seeds ca. 40–50, 1.8–4.8 mm long, 2.1–2.9 mm wide, 1.5–2.1 mm thick, obovate in outline, acute at both ends, reticulate-foveate with each face marked with ca. 12–17(-19) foveae. Germination epigeal.


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© Kristen Porter-Utley

Source: PhytoKeys

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