HabitTrailing or spreading perennial herbs, woody at the base, to 1 m tall, to 1 m or more in diameter. Stems 0.4-0.5 cm in diameter, pale grayish green, densely velvety pubescent with soft white, simple, uniseriate trichomes to 0.2 mm long from a unicellular base, these interspersed with longer uniseriate 3-4-celled trichomes from multicellular bases and tiny glandular trichomes with 4-celled glands (less often the glands 1 or 8-celled), the glandular trichomes more abundant on the young growth.
Sympodial StructureSympodial units 2-foliate; internodes 3-8 cm long.
LeavesLeaves interrupted imparipinnate, 5-12 cm long, 2-6 cm wide, dark green to pale green beneath, adaxially soft velvety pubescent with simple, uniseriate eglandular trichomes like those of the stems, occasionally with longer more robust trichomes from a darker, multicellular base, abaxially the pubescence similar but denser, evenly spread on the veins and the lamina, the abaxial surface paler due to more abundant pubescence; primary leaflets 2-3 pairs, the basal pair usually smaller, narrowly elliptic to elliptic, the base truncate, very oblique and decurrent basiscopically, the margins entire to deeply crenate, the crenations deeper near the base of the leaflets, the apex acute; terminal leaflet the same size or often larger than than the laterals, (1.5-) 2-4.8 cm long, (0.5-) 0.6-1.8 cm wide, the apex long acuminate, the petiolule 0.2-1 cm long, the apex acute; lateral leaflets (0.5-) 1-3.4 cm long, (0.3-) 0.4-1.5 cm wide, sessile or the petiolule to 0.5 cm long; secondary leaflets absent; tertiary leaflets absent; interjected leaflets 0-4, 0.1-0.9 cm long, 0.1-0.7 cm wide, sessile; petiole 1-3 cm long; pseudostipules present on some nodes, 0.5-1 cm long, 0.4-0.8 cm wide, the margin entire to irregularly crenate.
InflorescencesInflorescences (2-) 3-9 cm, simple or sometimes once branched, with 2-7 (-15) flowers, with bracts on the basal nodes, the bracts 0.3-0.7 cm long, 0.4-0.5 cm wide, peduncle (1-) 2-4 (-11) cm long, pubescent like the stems. Pedicels 0.7-1 cm long, articulated in the distal half. Buds 1-1.2 cm long, ca. 0.5 cm wide, conical, with the corolla only halfway exserted from the calyx lobes just before anthesis.
FlowersFlowers with the calyx tube 0.05-0.1 cm long, the lobes 0.5-0.7 cm long, 0.15-0.2 cm wide, lanceolate, sparsely to densely pubescent with small simple, uniseriate trichomes and a few glandular trichomes like those of the inflorescence axis; corolla 1.6-2 cm in diameter, rotate-stellate, bright yellow, the tube 0.3-0.5 cm long, the lobes 0.5-0.8 cm long, reflexed at anthesis; staminal column 0.9-1.1 cm long, straight, the filaments ca. 0.5 mm long, the anthers 0.6-0.7 cm long, the sterile apical appendage 0.15-0.2 cm long; ovary globose, or minutely papillate at the apex; style 0.8-0.9 cm long, ca. 0.5 mm in diameter, densely white pubescent in the basal half, straight, exserted 0.5-1 mm beyond the sterile appendage; stigma minutely capitate, green.
FruitsFruit 1-1.3 cm in diameter, globose, 2-locular, green, with a dark green stripe from the apex to the base, densely papillate and sparsely pubescent with extremely slender and weak simple uniseriate trichomes to 0.5 mm long; fruiting pedicels 1-1.3 cm long, straight or slightly angled at the articulation; calyx lobes in fruit 0.8-1 cm long, 0.2-0.3 cm wide, loosely investing the base of the berry to somewhat spreading.
SeedsSeeds 2.2-3.2 mm long, 1.2-1.6 mm wide, 0.6-0.8 mm thick, obovate, pale brown, pubescent with hair-like outgrowths of the lateral testa cell walls, these adpressed and giving a silky appearance to the surface, narrowly winged (ca. 0.2 mm) at the apex and acute at the base.
Life History and Behavior
Evolution and Systematics
Systematics or Phylogenetics
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