HabitShrubs or small trees, to 4 m tall, unarmed. Trunks to 12 cm in diameter; bark of older stems pale brown; young branches strongly sulcate, grayish to reddish, softly tomentose-pubescent, the indument persistent, becoming matted, the hairs short- to rather long-stalked, porrect-stellate, multangulate, dendritic-multangulate, and dendritic-echinoid, nearly furfuraceous.
Sympodial StructureSympodial units plurifoliate, axillary leaves lacking.
LeavesLeaves simple, the blades 14-32 x 4.2-12 cm, ca. 2.5 times as long as wide, ovate to ovate-elliptic, coriaceous, dark green, velutinous to glabrate adaxially, the hairs touching or lacking, short-stalked, porrect-stellate, whitish tomentose-pubescent abaxially, the hairs stalked, porrect-stellate; unarmed; main lateral veins 10-15 on each side of midrib; bases acute, obtuse, or cuneate; margins entire, occasionally appearing slightly crisped (wavy) in herbarium specimens; apex acute or attenuate; petioles 1-5 cm, ca. 1/14-1/6 the length of the blades, tomentose-pubescent, the hairs mixed sessile and stalked, porrect-stellate, multangulate, and dendritic-multangulate, nearly furfuraceous.
InflorescencesInflorescences 7-16 cm, pseudo-terminal, many times branched, with ca. 80-240 flowers, the axes unarmed, softly tomentose-pubescent, the hairs mixed sessile and stalked, porrect-stellate, multangulate, and dendritic-multangulate; peduncle (6.1-) 9.8-15.4 x 0.2-0.5 cm, sulcate; rachis 0.6-2.2 (-6.1) cm; pedicels 1.5-3 mm in flower, 4.5-5.5 mm in fruit, closely spaced 0.8-2.8 (-4.4) mm apart, articulated at the base.
FlowersBuds 6-8 mm long at anthesis, oblong; corolla tomentose-pubescent in bud, the hairs mixed sessile and short-stalked, porrect-stellate, thin-walled, transparent; calyx lobe sutures obscured by tomentum. Flowers all apparently perfect; calyx subcampanulate, 3.6-5.7 mm long, the tube 2.2-3.3 mm, lobed 1/2-2/3 way to the base, the lobes often splitting unevenly, 1.1-4 x 1.5-3 mm in flower, acute, obtuse or semi-ovate, chartaceous, somewhat translucent, tomentose-pubescent abaxially, the hairs sessile and short-stalked, porrect-stellate and multangulate, a few minute simple and porrect-stellate hairs adaxially on upper fourth of lobes; fruiting calyx accrescent, enveloping but not completely covering the fruit, pubescence as in flower, the lobes 2.9-6 x 5-6.5 mm. Corolla 1-2.2 (-3.1) cm in diameter, 8-14 mm long, soon exserted from calyx, nearly the length of calyx or its lobes by anthesis, stellate, chartaceous, white, the tube 2.3-3.8 mm, the lobes 6-10 x 2.7-4 (-5.5) mm, ovate to ovate-lanceolate, pubescence as in bud abaxially, with few simple and porrect-stellate hairs on upper fourth adaxially. Stamens with filaments 0.9-2 mm, inserted on corolla tube 0.9-1.9 mm above base, glabrous; anthers 3-4.6 x 0.8-1.5 mm, lanceolate, blunt-tipped, golden yellow, free, poricidal at the tips, the pores directed distally and somewhat adaxially, farinose around edges, lengthening to longitudinal slits with age. Ovary subconical to subglobose, tomentose-pubescent, the hairs porrect-stellate and multangulate; style 5.3-6.4 x 0.2-0.3 mm, exserted from stamens, cylindrical, densely pubescent on lower 2/3-3/4, the hairs porrect-stellate and multangulate; stigma capitate.
FruitsFruit a fleshy berry, 0.8-1.2 cm in diameter, globose, obtuse at apex, green, sparsely pubescent, the hairs sessile and short-stalked, porrect-stellate and multangulate.
SeedsSeeds 3.1-3.8 x 2.5-3 mm, suborbicular, yellowish-tan, the surface reticulate-punctate.
Habitat and Ecology
Evolution and Systematics
Systematics or Phylogenetics
IUCN Red List Assessment
Red List Category
Red List Criteria
- Needs updating
References and More Information