HabitProstrate, stoloniferous perennial shrub up to 0.3 m tall. Juvenile stage absent. Stems terete or ridged, brown, sparsely to densely stellate-pubescent, the stellae 0.2-0.35 mm in diameter, sessile, the lateral rays 7-8, porrect, the midpoints 0.5-1.5 times as long as lateral rays, eglandular; finger hairs absent; very short, gland-tipped hairs hairs absent; prickles straight, acicular, 3-7 mm long, 9-20 times longer than wide, 25-65 per dm.
Sympodial StructureSympodial units probably difoliate, the leaves geminate or not.
LeavesLeaves simple, the blades 5.5-9.5 x 2.8-4.5 cm, 1.6-2.3 times as long as wide, elliptical or ovate, chartaceous, green and glabrous to sparsely stellate-pubescent adaxially, the stellate hairs confined to midrib; finger hairs absent; very short, gland-tipped hairs absent; prickles present on midvein and lateral veins, 20-60, straight, acicular, 2-8 mm long; green to white or yellowish and sparsely to very densely stellate-pubescent abaxially, the hairs spaced 0.05-0.25 mm apart, 0.25-0.35 mm in diameter, sessile, the lateral rays 7-8, porrect, the midpoints 0.2-0.8 times as long as lateral rays, eglandular; finger hairs absent; very short, gland-tipped hairs absent; prickles present on midvein and lateral veins, 10-30, straight, acicular; base cuneate or obtuse, the oblique part 0-5 mm long; margin entire or shallowly lobed throughout, the lobes 2-4 on each side, acute, lobing index 1-1.4; apex acute; petioles 1-1.7 cm long, 14-29% length of lamina, sparsely to densely stellate-pubescent, prickles present.
InflorescencesInflorescence up to 1 cm long, extra-axillary, unbranched, pseudo-umbellate or pseudo-racemose, with 2-8 flowers, the plants strongly or weakly andromonoecious, the axes sparsely to densely stellate-pubescent, armed or unarmed; peduncle 0-0.7 cm; rachis 0.2-0.3 cm; pedicels 6-20 mm long in flower, 0.25-0.4 mm thick throughout, 21-30 mm long in fruit, ca. 0.8 mm thick, spaced 1-2 mm apart, articulated at the base.
FlowersFlowers heterostylous, 5-merous. Calyx 5-6 mm long, the tube 2-3 mm long, the lobes 1.5-3.5 mm long, attenuate, moderately stellate-pubescent, the hairs transparent or purple, 0.2-0.3 mm across, sessile, the lateral rays 7-8, the midpoints 0.5-1 times as long as laterals, eglandular; finger hairs absent; very short, gland-tipped hairs absent; prickles present or absent, 0-5 per flower, 0.5-2 mm long; fruiting calyx with lobes less than half length of mature fruit, armed or unarmed. Corolla 1.5-2.5 cm in diameter, 8-12 mm long, shallowly lobed, with sparse interpetalar tissue, chartaceous, pinkish or purple, the tube 3-5 mm, the lobes 9-10 x 4-6 mm, narrowly triangular, moderately stellate-pubescent abaxially on center of lobes, glabrous adaxially and on interpetalar tissue. Stamens with filaments 0.5-1.5 mm; anthers 4-5 x 1-1.5 mm at base, narrowly ovate, slightly tapered, not or loosely connivent, yellow, the pores minute and directed distally. Ovary with very short, gland-tipped hairs on distal half; functional style 6-7.5 x 0.5-1 mm, cylindrical, glabrous or with very short, gland-tipped hairs at base; stigma capitate.
FruitsFruits 1 per inflorescence, 1.5-1.8 cm in diameter, 1-locular (septum absent or incomplete), globose, yellowish-green, glabrous, the exocarp ca. 0.8 mm thick, the mesocarp moist but not juicy, the placenta in cross-section sessile, elliptical.
SeedsSeeds 3.5-4 x 3-3.5 mm, whitish or pale yellow, flattened, the surface finely reticulate.
Life History and Behavior
Evolution and Systematics
Systematics or Phylogenetics
References and More Information