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Species of subgenus Apocephalus are all endoparasitoids of ants (Hymenoptera: Formicidae), including the type species, which is the original ""ant-decapitating fly."" A diversity of lifestyles exist within the ant parasitoids, but most are solitary larvae developing in the ant head. Subgenus Mesophora are parasitoids of stingless bees, bumble bees, spiders, and especially cantharoid beetles (Cantharidae, Lampyridae). The way of life of most species is unknown.