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Heteroponera Mayr Reinstated HNS (Hymenoptera: Formicidae HNS ).

- Heteroponera Mayr HNS (1887, Verh. zool.-bot. Ges. Wien, 37: 533) has remained suppressed as a synonym of Acanthoponera Mayr HNS for many years. Wheeler (1923, loc. cit.) has shown, however that Acanthoponera HNS can be split into two groups; one group { Acanthoponera HNS s. str.) having the tarsal claws with an extra, strong tooth and a basal lobe (or tooth), while the second group (Anacanthoponera Wheeler) has the tarsal claws at most with a single, weak median tooth. In this second group, Wheeler included Heteroponera carinifrons Mayr HNS , and since Heteroponera HNS , with the genotype H. carinifrons HNS , has precedence, it must be reinstated as a good genus. Anacanthoponera Wheeler (1923, Psyche, 30: 176, as a subgenus of Acanthoponera HNS ) is a new synonym of Heteroponera HNS , since the genotype, Ponera dolo Roger HNS , is congeneric with H. carinifrons HNS .

Acanthoponera HNS is neotropical, and has well developed propodeal teeth and the petiolar apex produced as a long tooth or spine; Kusnezov (in litt.) finds A. mucronata HNS to have 6, 4 palpal segmentation. Heteroponera HNS is neotropical and Australasian, and the propodeal teeth and dentiform petiolar apex are absent or feebly developed; H. imbellis Emery HNS has 3, 3 palpal segmentation (my dissection). Until more species can be critically examined, Acanthoponera HNS and Heteroponera HNS should be considered as distinct genera. Acanthoponera HNS appears to be the most generalized living member of the Ectatommini HNS , and is probably close to the stem from which the proceratiines, the myrmicines and Paraponera HNS arose. - W. L. Brown, Jr., Museum of Comparative Zoology, Harvard University.


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