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DescriptionAnnual, biennial or perennial herbs. Leaves alternate. Inflorescence of scorpioid cymes. Flowers bisexual. Calyx 5-lobed, not or only slightly accrescent. Corolla tube short, cylindric or funnel-shaped, closed with 5 scales at the throat; lobes 5. Ovary of 4 separate lobes. Style gynobasic. Fruit of 4 nutlets; nutlets depressed, glochidiate.