Overview

Brief Summary

Introduction

Often treated as N. candida var. copelandii, Notholaena copelandii differs from N. candida in morphology (Hall, 1950), farina chemistry (Wollenweber, 1984; Wollenweber et. al 2001), geographic range, and plastid DNA sequences (Rothfels et al., 2008). I thus follow Windham (1993a) and Mickel and Smith (2004) in treating them as distinct species.

Notholaena copelandii
is a species of southern Texas and northeastern Mexico, where it occurs on limestone cliffs and outcrops.

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Comprehensive Description

Characteristics

The morphological characters distinguishing N. copelandii from N. candida include: basal pinnae shape (a single exaggerated basioscopic pinnule on N. copelandii; pinnules gradually tapering in length in N. candida); leaf tip shape (abruptly tapered vs. gradually tapered); and rhizome scales (ciliate vs. slightly toothed). Notholaena copelandii is also similar to N. jacalenis, but the later species has much narrower leaves.

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Distribution

National Distribution

United States

Origin: Unknown/Undetermined

Regularity: Regularly occurring

Currently: Unknown/Undetermined

Confidence: Confident

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Tex.; Mexico.
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Physical Description

Morphology

Description

Stem scales strongly bicolored, margins brown, broad and well defined, thin, weakly ciliate. Leaves 6--25 cm. Petiole black, equal to or somewhat longer than blade, rounded adaxially, glabrous except for a few scales and farinose glands near base. Blade ovate, 2-pinnate-pinnatifid, 1--2 times longer than wide, abaxially with conspicuous white farina, scales absent, adaxially glandular; basal pinnae much larger than adjacent pair, strongly inequilateral, proximal basiscopic pinnules greatly enlarged. Ultimate segments sessile, broadly adnate to costae; segment margins slightly recurved, rarely concealing sporangia. Sporangia containing 64 spores. 2 n = 60.
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Diagnostic Description

Synonym

Cheilanthes candida M. Martens & Galeotti var. copelandii (C. C. Hall) Mickel; Chrysochosma candida (M. Martens & Galeotti) Kümmerle var. copelandii (C. C. Hall) Pichi-Sermolli; Notholaena candida (M. Martens & Galeotti) Hooker var. copelandii (C. C. Hall) R. M. Tryon
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Ecology

Habitat

Sporulating summer--fall. Rocky slopes and cliffs, apparently confined to limestone; 300--1500m.
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Conservation

Conservation Status

National NatureServe Conservation Status

United States

Rounded National Status Rank: NNR - Unranked

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NatureServe Conservation Status

Rounded Global Status Rank: G4 - Apparently Secure

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Notes

Comments

Notholaena copelandii often has been treated as a variety of the Mexican species N . candida (M. Martens & Galeotti) Hooker. The two taxa show significant differences in stem scale and leaf morphology (R. M. Tryon 1956), density of glands on the adaxial leaf surface, and chemical composition of the farinose indument (E. Wollenweber 1984). Measures of genetic similarity based on isozyme data indicate that N . copelandii and N . candida are quite distinct and, at present, allopatry precludes gene flow. Past hybridization may account for the small number of intermediate specimens reported from San Luis Potosí, Mexico, by R. M. Tryon (1956).
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