The adults are gnats that consume other small insects or bite warm-blooded animals. In addition to these activities, they also visit flowers with exposed nectaries, but aren't important pollinators. The larvae live in moist ground where there is a suitable amount of decaying organic material.
Small flies, 1-6 mm long, slender to moderately robust. Female with biting and sucking mouthparts; bloodsucker of vertebrates or insects, or predator. Male with plumose antennae. Wings held horizontally at rest one above the other, sometimes strongly patterned. Early stages usually found in moist or aquatic habitats; larva apneustic, often elongate in form, and a strong swimmer. (Downes & Wirth 1981)
Known prey organisms
Based on studies in:
New Zealand: Otago, Berwick, Meggatburn (River)
New Zealand: South Island, Canton Creek, Taieri River, Lee catchment (River)
New Zealand: Otago, Stony, Sutton catchment (River)
This list may not be complete but is based on published studies.
- Thompson, RM and Townsend CR. 2005. Energy availability, spatial heterogeneity and ecosystem size predict food-web structure in streams. OIKOS 108: 137-148.
- Townsend, CR, Thompson, RM, McIntosh, AR, Kilroy, C, Edwards, ED, Scarsbrook, MR. 1998. Disturbance, resource supply and food-web architecture in streams. Ecology Letters 1:200-209.
Molecular Biology and Genetics
Statistics of barcoding coverage
Specimens with Sequences:24742
Specimens with Barcodes:22947
Species With Barcodes:364
Ceratopogonidae, or biting midges are a family of small flies (1–4 mm long) in the order Diptera. They are also known as no-see-ums, midgies, sand flies, punkies, and others in North America, and sandflies in Australia. They are closely related to the Chironomidae, Simuliidae (or black flies), and Thaumaleidae.
They are found in almost any aquatic or semiaquatic habitat throughout the world, as well as in mountain areas. Females of most species are adapted to suck blood from some kind of host animal. Culicoides, Forcipomyia (Lasiohelea), and Leptoconops suck vertebrate blood. Some Atrichopogon and Forcipomyia species are ectoparasites on larger insects. Dasyhelea species feed exclusively on nectar. Species in other genera are predatory on other small insects. Larvae are always found in some damp location, such as under bark, in rotten wood, compost, mud, stream margins, tree holes, or water-holding plants (i.e., phytotelmata).
Many of the hematophagic (blood-eating) species are pests in beach or mountain habitats. Some other species are important pollinators of tropical crops such as cacao. The blood-sucking species may be vectors of disease-causing viruses, protozoa, or filarial worms. The bite of midges in the genus Culicoides causes an allergic response in equines known as sweet itch. In humans, their bites can cause intensely itchy, red welts that can persist for more than a week. The discomfort arises from a localized allergic reaction to the proteins in their saliva, which can be somewhat alleviated by topical antihistamines.
The smaller members of the family are tiny enough to pass through the apertures in typical window screens. Camping tents are often equipped with extra-fine mesh netting, called no-see-um nets, to keep the pests out.
- Blanton, F.S. and W.W. Wirth. 1979. The sand flies (Culicoides) of Florida (Ceratopogonidae). Arthropods of Florida and Neighboring Land Areas Volume 10. Florida Department of Agriculture and Consumer Services.
- Borkent, A. and W.W. Wirth. 1997. World species of biting midges (Diptera: Ceratopogonidae). Bulletin of the American Museum of Natural History 233: 1–257.
- Clastrier, J. and W.W. Wirth. 1978. The Leptoconops kerteszi complex in North America (Diptera: Ceratopogonidae). United States Department of Agriculture Technical Bulletin Number 1573.
- Downes, J.A. and W.W. Wirth. 1981. Chapter 28: Ceratopogonidae. Pp. 393–421. In: McAlpine, J.F., B.V. Peterson, G.E. Shewell, H.J. Teskey, J.R. Vockeroth, and D.M. Wood. Manual of Nearctic Diptera, Volume 1. Agriculture Canada Monograph 27.
- Hendry, George. Midges in Scotland 4th Edition, Mercat Press, Edinburgh, 2003 ISBN 1-84183-062-3
- Mullen, G.R. and L.J. Hribar. 1988. Biology and feeding behavior of ceratopogonid larvae (Diptera: Ceratopogonidae) in North America. Bulletin of the Society for Vector Ecology 13: 60–81.
- Wirth, W.W. and F.S. Blanton. 1974. The West Indian sandflies of the genus Culicoides (Diptera: Ceratopogonidae). United States Department of Agriculture Technical Bulletin Number 1474.
- Wirth, W.W. and W.L. Grogan, Jr. 1988. The Predaceous Midges of the World (Diptera: Ceratopogonidae; Tribe Ceratopogonini). Flora and Fauna Handbook Number 4. E.J. Brill Publishers, Leiden. xv + 160 pp.
- Wirth, W.W., N.C. Ratanaworabhan, and D.H. Messersmith. 1977. Natural history of Plummers Island, Maryland. XXII. Biting midges (Diptera: Ceratopogonidae). 1. Introduction and key to genera. Proceedings of the Biological Society of Washington, 90(3): 615–647.
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