Regularity: Regularly occurring
California Central Valley Grasslands Habitat
This taxon is found in the California Central Valley grasslands, which extend approximately 430 miles in central California, paralleling the Sierra Nevada Range to the east and the coastal ranges to the west (averaging 75 miles in longitudinal extent), and stopping abruptly at the Tehachapi Range in the south. Two rivers flow from opposite ends and join around the middle of the valley to form the extensive Sacramento-San Joaquin Delta that flows into San Francisco Bay.
Perennial grasses that were adapted to cool-season growth once dominated the ecoregion. The deep-rooted Purple Needle Grass (Nassella pulchra) was particularly important, although Nodding Needle Grass (Stipa cernua), Wild Ryes (Elymus spp.), Lassen County Bluegrass (Poa limosa), Aristida spp., Crested Hair-grass (Koeleria pyramidata), Deergrass (Muhlenbergia rigens,), and Coast Range Melicgrass (Melica imperfecta) occurred in varying proportions. Most grass growth occurred in the late spring after winter rains and the onset of warmer and sunnier days. Interspersed among the bunchgrasses were a rich array of annual and perennial grasses and forbs, the latter creating extraordinary flowering displays during certain years. Some extensive mass flowerings of the California Poppy (Eschscholzia californica), Lupines (Lupinus spp.), and Exserted Indian Paintbrush (Castilleja exserta) are found in this grassland ecoregion.
Prehistoric grasslands here supported several herbivores including Pronghorn Antelope (Antilocapra americana), elk (including a valley subspecies, the Tule Elk, (Cervus elaphus nannodes), Mule Deer (Odocoileus hemionus), California ground squirrels, gophers, mice, hare, rabbits, and kangaroo rats. Several rodents are endemics or near-endemics to southern valley habitats including the Fresno Kangaroo Rat (Dipodomys nitratoides exilis), Tipton Kangaroo Rat (Dipodomys nitratoides nitratoides), San Joaquin Pocket Mouse (Perognathus inornatus), and Giant Kangaroo Rat (Dipodomys ingens). Predators originally included grizzly bear, gray wolf, coyote, mountain lion, ringtail, bobcat, and the San Joaquin Valley Kit Fox (Vulpes velox), a near-endemic.
The valley and associated delta once supported enormous populations of wintering waterfowl in extensive freshwater marshes. Riparian woodlands acted as important migratory pathways and breeding areas for many neotropical migratory birds. Three species of bird are largely endemic to the Central Valley, surrounding foothills, and portions of the southern coast ranges, namely, the Yellow-billed Magpie (Pica nuttalli), the Tri-colored Blackbird (Agelaius tricolor EN), and Nuttall’s Woodpecker (Picoides nuttallii).
The valley contains a number of reptile species including several endemic or near-endemic species or subspecies such as the San Joaquin Coachwhip (Masticophis flagellum ruddocki), the Blunt-nosed Leopard Lizard (Gambelia sila EN), Gilbert’s Skink (Plestiodon gilberti) and the Sierra Garter Snake (Thamnophis couchii). Lizards present in the ecoregion include: Coast Horned Lizard (Phrynosoma coronatum NT); Western Fence Lizard (Sceloporus occidentalis); Southern Alligator Lizard (Elgaria multicarinata); and the Northern Alligator Lizard (Elgaria coerulea).
There are only a few amphibian species present in the California Central Valley grasslands ecoregion. Special status anuran taxa found here are: Foothill Yellow-legged Frog (Rana boylii NT); Pacific Chorus Frog (Pseudacris regilla); and Western Spadefoot Toad (Pelobates cultripes). The Tiger Salamander (Ambystoma tigrinum) occurs within this ecoregion.
Although many endemic plant species are recognized, especially those associated with vernal pools, e.g. Prickly Spiralgrass (Tuctoria mucronata). A number of invertebrates are known to be restricted to California Central Valley habitats. These include the Delta Green Ground Beetle (Elaphrus viridis CR) known only from a single vernal pool site, and the Valley Elderberry Longhorn Beetle (Desmocerus californicus dimorphus) found only in riparian woodlands of three California counties.
Vernal pool communities occur throughout the Central Valley in seasonally flooded depressions. Several types are recognized including valley pools in basin areas which are typically alkaline or saline, terrace pools on ancient flood terraces of higher ground, and pools on volcanic soils. Vernal pool vegetation is ancient and unique with many habitat and local endemic species. During wet springs, the rims of the pools are encircled by flowers that change in composition as the water recedes. Several aquatic invertebrates are restricted to these unique habitats including a species of fairy shrimp and tadpole shrimp.
- World Wildlife Fund & C. Michael Hogan. 2013."California Central Valley grasslands". Encyclopedia of Earth, National Council for Science and the Environment, Washington DC ed.Mark McGinley.
- Michael G.Barbour, Todd Keeler-Wolf and Allan A. Schoenherr. 2007. Terrestrial vegetation of California. 712 pages
National NatureServe Conservation Status
Rounded National Status Rank: NNR - Unranked
Castilleja exserta (formerly Orthocarpus purpurascens) is a species of plant in the genus which includes the Indian paintbrushes. Its common names include purple owl's clover, escobita, and exserted Indian paintbrush. 
The Castilleja exserta plant is an annual about 1 foot (0.30 m) tall with a hairy stem covered in thready leaves. Like other related Castilleja plants, this is a hemiparasite which derives some of its nutrients directly from the roots of other plants by injecting them with haustoria. This is the reason for its small, reduced leaves.
Although this species is variable in appearance and easily hybridizes with other Castilleja species, it generally bears a brightly colored inflorescence of shaggy pink-purple or lavender flowers. The thin, erect bracts are usually tipped with the same color, giving the inflorescence the appearance of a paintbrush.
- Castilleja exserta ssp. exserta — pale purple owlclover. 
- Castilleja exserta ssp. latifolia — wideleaf Indian paintbrush; endemic to coastal California. 
- Castilleja exserta ssp. venusta — endemic to California: Mojave Desert, southern San Joaquin Valley.  
Bay checkerspot butterfly host
- CalFlora database: Castilleja exserta . accessed 3.23.2013
- USDA: C. exserta . accessed 3.23.2013
- CalFlora: Castilleja exserta ssp. exserta
- CalFlora: Castilleja exserta ssp. latifolia
- CalFlora: Castilleja exserta ssp. venusta
- USDA: C. e. ssp. venusta
- Lady Bird Johnson Wildflower Center Native Plant Information Network (NPIN) — Castilleja exserta (Purple owl's clover) . accessed 3.23.2013
Names and Taxonomy
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