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 Perennial herbs, 10–40 cm tall. Stems erect, inconspicuously ribbed, puberulous. Leaves simple, rosulate or alternate at base, 5–15 by 1.5–5 cm, obovate or oblanceolate, margin slightly serrate to entire, apex obtuse or rounded, base cuneate or attenuate, subcoriaceous; upper surface puberulous without glands, lower surface puberulous with filiform hairs and capitate glands; lateral veins 9–15-paired; petioles up to 2 cm long. Capitulescences terminal and axillary, capitula 1-4 aggregated in clusters supported by foliaceous bracts, clusters arranged in a spike. Capitula tubular, 14–17 mm long. Receptacle flat, 1–1.5 mm in diam., glabrous. Florets bisexual and fertile. Involucres oblong, in 2 series, 10–11 mm long. Phyllaries 8, decussate, light green, margin entire or piliferous, outer surface pilose without glands; the outer lanceolate, apicies acute; the inner ones obovate-lanceolate or oblong, apices acute. Florets 4; corollas salverform, white, zygomorphic, glabrous; corolla tubes 5–9 mm long; corolla lobes 2.5–2.8 mm long. Anthers 1.5–2 mm long, apical appendage acute, base acute. Styles white, 5–9 mm long, branches ca. 2 mm long, inner surface covered with stigmatic papillae. Achenes clavate, 4–5 mm long, pubescent with densely twin hairs, without glands, 10-ribbed. Pappus in 1 series, often of 2 sizes and bent at the tip, bristles 6–9, 2–6 mm long.


Creative Commons Attribution 3.0 (CC BY 3.0)

© Sukhonthip Bunwong, Pranom Chantaranothai, Sterling C. Keeley

Source: PhytoKeys

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