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Perennials, 50–200+ cm (rhizomatous, producing tubers late in growing season). Stems erect, scabro-hispid to hirsute (sometimes glaucous). Leaves mostly cauline; opposite or alternate proximally, usually alternate distally; petioles 2–8 cm (often ± winged); blades (3-nerved from near bases) lanceolate to ovate, 10–23 × 7–15 cm, bases broadly to narrowly cuneate, margins entire or serrate (flat), abaxial faces puberulent or hirsutulous to tomentulose and gland-dotted (abaxial) or ± scabrous (adaxial). Heads 3–15. Peduncles 1–15 cm. Involucres hemispheric, (10–25 ×) 8–12 mm. Phyllaries (often dark green, drying nearly black) 22–35 (bases appressed, apices ± spreading, sometimes reflexed in fruit), lanceolate, 8.5–15 × 2–4 mm (subequal), (margins ciliate) apices acuminate, abaxial faces hispidulous or puberulent, gland-dotted. Paleae 8–9 mm, 3-toothed (apices hairy). Ray florets 10–20; laminae 25–40 mm. Disc florets 60+; corollas 6–7 mm, lobes yellow; anthers dark brown or black, appendages dark or yellowish. Cypselae 5–7 mm, glabrous or distally hairy; pappi of 2 aristate scales 1.9–3 mm plus 0–1 deltate scales 0.5–0.8 mm. 2n = 102.


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© Missouri Botanical Garden, 4344 Shaw Boulevard, St. Louis, MO, 63110 USA

Source: Missouri Botanical Garden


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