Comprehensive Description

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Tetramorium insolens (F. Smith) HNS

(Figs 40, 45)

Myrmica insolens F. Smith HNS , 1861: 47. Holotype female, Sulawesi: Menado (A. R. Wallace) (UM, Oxford) [examined]. Tetramorium insolens (F. Smith) HNS ; Donisthorpe, 1932: 468. [For a full statement of current synonymy of insolens HNS see Bolton, 1977: 99 with the exception of melanogyna Mann HNS , for which see p. 173, this paper.]

Worker. TL 3.3 - 4.0, HL 0.78 - 0.94, HW 0.68 - 0.84, CI 84 - 88, SL 0.56 - 0.68, SI 78 - 86, PW 0.50 - 0.62, AL 0.92 - 1.08 (40 measured).

Figs 41 - 49 Tetramorium HNS workers. Head or alitrunk of (41) simillimum HNS , (42) caldarium HNS , (43) bicarinatum HNS , (44) pacificum HNS , (45) insolens HNS , (46) placidum HNS , (47) bicarinatum HNS , (48) lucayanum HNS , (49) caespitum HNS . Pilosity omitted in Figs 47 - 49.

Figs 50 - 59 Tetramorium HNS workers. Head and alitrunk of (50, 51) hispidum HNS , (52, 53) mexicanum HNS . 54 - 55. Hind tibia of (54) mexicanum HNS , (55) spinosum HNS to show pilosity. 56 - 59. Alitrunk of (56) palaense HNS , (57) ocothrum HNS , (58) rekhefe HNS , (59) belgaense HNS . Fringing pilosity only indicated in Figs 50 and 52.

Mandibles smooth and shining, unsculptured except for scattered pits. Anterior clypeal margin with a median notch or impression; median portion of clypeus with three strong longitudinal carinae. Frontal carinae long and strong, extending back almost to occiput. Eyes of moderate size, maximum diameter c. 0.18 - 0.20, about 0.23 - 0.26 x HW. Pronotal corners in dorsal view angular. Propodeal spines long and stout, generally somewhat upcurved along their length, more rarely with the extreme apex of each spine suddenly upcurved. Metapleural lobes triangular, acute, somewhat upcurved. Petiole node in profile with anterior and posterior faces approximately parallel, the dorsum convex and rising slightly posteriorly so that the anterodorsal angle is on a slightly lower level than the posterodorsal, the latter angle usually sharper than the former, which has a tendency to be rounded. Dorsum of head to level of eyes with sparse longitudinal rugulae with a few cross-meshes and a fine but fairly conspicuous groundsculpture. Behind the level of the eyes the head is reticulate-rugose. Dorsal alitrunk with an irregular rugoreticulum which is usually strongest on the pronotum. Petiole and postpetiole reticulate-rugose both laterally and dorsally. Gaster unsculptured. All dorsal surfaces of head and body with abundant long erect or suberect hairs, those situated in a row on the upper surfaces of the frontal carinae between antennal insertion and occiput very long, distinctly longer than the maximum diameter of the eye. Colour varying from clear pale yellow to light orange-brown, usually with the gaster distinctly lighter in shade than the head and alitrunk. More rarely the ant is uniformly coloured.

The differences between this species and the closely related bicarinatum HNS were tabulated in part 2 (Bolton, 1977) of this study. Basically, insolens HNS differs from bicarinatum HNS in having smooth mandibles, relatively long hairs on the frontal carinae (longer than maximum diameter of eye), gaster not darker in colour than alitrunk and the petiole a slightly different shape (compare Figs 43 and 45).

As a tramp-species insolens HNS is by no means as successful as bicarinatum HNS . It has a very wide range in the Pacific island-systems and is known from Indonesia, Philippines and the Solomon Islands. It was collected by E. O. Wilson in Sri Lanka, where it may be an introduction, and in England (in orchid house), Germany (zoological garden) and in the U. S. A. (near Los Angeles) where it is most certainly introduced. The Los Angeles specimens are deposited in USNM, Washington; for other localities see Bolton (1977).


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