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Range DescriptionThis species is endemic to a narrow belt of subalpine meadows in the Great Caucasus Range of Georgia and Russia. Populations have been recorded from Mount Elbrus and the Baksan River in the east to the Fisht and Osten mountains in the west. Further to the west it is widely distributed in the upper reaches of the Kuban', Teberda, Zelenchuk, Belaya, Laba rivers and other waterbodies flowing to the north. Populations have also been recorded from the southern slopes of the Caucasus, in the upper reaches of the Bzyb' River and its tributaries in Abkchazia and in the upper reaches of the Mzymta River from the slopes of Mount Aibga. It has been recorded between 1,650 and 2,800m asl.