Range Description

This species was originally found in Quebec and New Brunswick west to Alberta and south to Wyoming, Arkansas, and Virginia (Burch, 1989); western occurrences are strongly disjunct and specimens from the Washington and Montana populations need to be compared in detail with more easterly occurrences, in view of the fact that speciation has occurred in several other genera with disjunct species swarms with both eastern and western representation. Recently, the species has been found in several lakes in the Clark Fork and Flathead drainages, Lake Roosevelt, Washington, Ferry Co., the Flathead Indian Reservation, Montana, searches in Idaho turned up no sites (Frest and Johannes 1995).