Distribution: East shore of Caspian Sea in Turkmenistan eastward to E Kazakhstan, Kyrgyzstan, Tajikistan, Uzbekistan, E Turkmenistan N Iran, Afghanistan, W Pakistan, W China (Xinjiang through Gansu to W Nei Mongol = Inner Mongolia and Ningxia), S Mongolia speciosus: Pakistan (KHAN 2002, pers. comm.); Tajikistan (Vakhsh River valley);
Type locality: “Eastern Bukhara” (Vakhsh River valley in Tajikistan)
Type locality: Tataria
Type locality: coast of the Aral Sea (fide KHALIKOV, pers. comm.)
Life History and Behavior
Lifespan, longevity, and ageing
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