Molecular Biology and Genetics
Statistics of barcoding coverage
Specimens with Sequences:598
Specimens with Barcodes:420
Species With Barcodes:72
Cannabaceae is a small family of flowering plants. As now circumscribed, the family includes about 170 species grouped in about 11 genera, including Cannabis (hemp), Humulus (hops) and Celtis (hackberries). Celtis is by far the largest genus, containing about 100 species.
Cannabaceae are often dioecious (distinct male and female plants). The flowers are actinomorphic (radially symmetrical) and not showy, as these plants are pollinated by the wind. As an adaptation to this kind of pollination, the calyx is short and there is no corolla. Flowers are grouped to form cymes. In the dioecious plants the masculine inflorescences are long and look like panicles, while the feminine are shorter and bear less flowers. The pistil is made of two connate carpels, the usually superior ovary is unilocular; there is no fixed number of stamens.
Classification systems developed prior to the 1990s, such as those of Cronquist (1981) and Dahlgren (1989), typically recognized the order Urticales, which included the families Cannabaceae, Cecropiaceae, Celtidaceae, Moraceae, Ulmaceae and Urticaceae, as then circumscribed. Molecular data from 1990s onwards showed that these families were actually embedded within the order Rosales, so that from the first classification by the Angiosperm Phylogeny Group in 1998, they were placed in an expanded Rosales, forming a group which has been called "urticalean rosids".
A molecular phylogenetic study in 2002 produced the consensus tree shown below. This showed that the family Celtidaceae was paraphyletic if the Cannabaceae, as then circumscribed, were removed. Accordingly a single larger family was required, combining the two existing families. Under the rules of the International Code of Nomenclature for algae, fungi, and plants, the oldest family name must be used: this is Cannabaceae.
List of genera
- Aphananthe Planchon (syn. Mirandaceltis Sharp)
- Cannabis L. – Hemp
- Celtis L. (syn. Sparrea Hunz. & Dottori)
- Gironniera Gaudich. (syn. Helminthospermum Thwaites, Nematostigma Planchon)
- Humulus L. (syn. Humulopsis Grudz.) – Hop
- Lozanella Greenman
- Parasponia Miquel
- Pteroceltis Maxim.
- Trema Loureiro (syn. Sponia Decaisne)
Carbon dating has revealed that these plants may have been used for ritual/medicinal purposes in Xinjiang, China as early as 494 B.C.
Hop (Humulus lupulus) is cultivated for its flowers which contain aromatic substances used in the production of beer. Its young shoots are used as vegetable. Different subspecies of hemp (Cannabis sativa) are cultivated for the production of fiber, as a source of cheap oil, for the nutritious seeds, or to produce recreational, sacramental, or medicinal cannabis.
- Stevens, P.F. (2001 onwards) "Cannabaceae", Angiosperm Phylogeny Website, retrieved 2014-02-25
- Sytsma, Kenneth J.; Morawetz, Jeffery; Pires, J. Chris; Nepokroeff, Molly; Conti, Elena; Zjhra, Michelle; Hall, Jocelyn C. & Chase, Mark W. (2002), "Urticalean rosids: circumscription, rosid ancestry, and phylogenetics based on rbcL, trnL-F, and ndhF sequences", American Journal of Botany 89 (9): 1531–1546, doi:10.3732/ajb.89.9.1531, PMID 21665755
- Stevens, P.F. (2001 onwards) "Cannabaceae Genera", Angiosperm Phylogeny Website, retrieved 2014-02-25
- Jiang, Hong-En; Xiao Li, You-Xing Zhao, David K. Ferguson, Francis Hueber, Subir Bera, Yu-Fei Wang, Liang-Cheng Zhao, Chang-Jiang Liu, Cheng-Sin Li (December 2006), "A new insight into Cannabis sativa (Cannabaceae) utilization from 2500-year-old Yanghai Tombs, Xinjiang, China", Journal of Ethnopharmacology 108 (3): 414–422, doi:10.1016/j.jep.2006.05.034, PMID 16879937, retrieved 8 June 2012
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