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Arthropleura is a genus of massive millipede-like terrestrial arthropods (segmented body, exoskeletal 'shell', and no internal bones). The largest of these extinct giants reached more than 2 meters in length and more than 46 centimeters in width (Lockley & Meyer 2000, Briggs et al. 1984, Martino & Grebe 2009). It is currently considered the largest terrestrial invertebrate ever. Arthropleurids lived in the Late Carboniferous (340- 280 million years ago), and died off at the beginning of the Permian (Holmes 2008). They were very widespread, ranging throughout North America and Europe (Wilson 2003). As detritivores, Arthropleura fed on fallen leaves, stems, and other decaying organic matter.