Bombyx mori's preferred food is white mulberry leaves, but it may also eat the leaves of any other mulberry tree (e.g., Morus rubra or Morus nigra) as well as the Osage Orange. Silkworm larvae grow to about 4 cm long, and then build a cocoon in which to pupate. This cocoon is collected and boiled to harvest the 300-900 meter long single silk thread from which it is made. Sometimes the boiled pupae are then eaten (called ground cucumber). Like most adults in the family Bombycidae, B. mori moths have reduced mouth parts so do not feed. Male and female moths are similarly colored, but where males have a wingspan of 3–5 cm, females have vestigial wings and much larger bodies, holding hundreds of eggs.
Bombyx mori has been used as a model organism in biological and genetic studies, and its full genome (~432 Mb) was sequenced and published in 2008.
- Wikipedia, The Free Encyclopedia, 9 September, 2011. "Bombyx mori". Retrieved 9 September 2011, from http://en.wikipedia.org/w/index.php?title=Bombyx_mori&oldid=449326047.
Bombyx mori originally existed in the wild throughout Asia. Though they are believed to no longer exist in the wild, they are in the care of the silk industry in Asia and Australia (Savela 1998).
Biogeographic Regions: oriental (Native ); australian (Native )
The larvae of B. mori are caterpillars that are about 4 cm long, including their horned tail. They are buff-colored with brown thoracic markings. The adults are moths with a 4 cm wingspan. They are also buff-colored, but have thin brown lines on their whole bodies (Herbison-Evans 1997). Another silkworm, Bombyx mandarina, appears to be a wild race of B. mori (Savela 1998).
Although B. mori is native to China, it does not live in the wild any longer because of sericulture (Encarta 1998).
Terrestrial Biomes: forest
Bombyx mori are herbivores. They feed specifically on white mulberry leaves, but also eat Osage oranges and lettuce. They do most of their eating in the larval stage (Encarta 1998). The larvae have mandibles for feeding, while the adults have sucking mouth parts (Lepidoptera Part 2 1997). Because they have been cultivated for so long for sericulture (the silk industry), B. mori have lost an adaptation helpful to feeding in the wild. The larvae can no longer hang on plants at gravity-defying angles, and must be fed by humans (Herbison-Evans 1997).
Life History and Behavior
Bombyx mori are holometabolous and reproduce sexually. The female adult dies upon depositing her eggs (Encarta 1998). These eggs weigh in at a miniscule 1/30,000 of an ounce each (Knowledge Adventure 1997). After 10 days, the eggs hatch and hungry larvae emerge. They are segmented and have body hair. The larvae eat and grow for approximately 6 weeks, and then they begin the next stage of their lives. Bombyx mori produce a fluid in their silk glands that is forces through spinnerets on their mouths. This fluid hardens in the air to produce the silk thread that they will wrap around themselves to form their cocoons. Bombyx mori spend 2 weeks as pupae in the safety of their cocoons before emerging as adults (Encarta 1998). Inside the cocoon, much of their bodies die by an attack of their own digestive juices. This process, histolysis, clears away the old parts to make way for the new ones that will develop in this pupal state. After this process is completed, the adults break free from the cocoon in order to begin the cycle again. The adults are winged and have traded body hair for scales. They are dramatically different form their larval stage (Lepidoptera Part 2 1997).
Evolution and Systematics
The antennae of male silk moths distribute chemicals by having an oily coating that serves a dual purpose by binding to lipid-loving female pheromones and transporting them to nerve cells.
"[A] coating on the male silk moth's antenna...helps it smell nearby female moths. The coating catches pheromone molecules in the air and carries them through nanotunnels in the exoskeleton to nerve cells that send a message to the bug's brain. 'These pheromones are lipophilic. They like to bind to lipids, or fat-like materials. So they get trapped and concentrated on the surface of this lipid layer in the silk moth. The layer greases the movement of the pheromones to the place where they need to be,' [according to Michael Mayer, an associate professor in the University of Michigan's departments of Biomedical Engineering and Chemical Engineering]" (Moore 2011:1)
Learn more about this functional adaptation.
- Yusko EC; Johnson JM; Majd S; Prangkio P; Rollings RC; Jiali L; Yang J; Mayer M. 2011. Controlling protein translocation through nanopores with bio-inspired fluid walls. Nature Nanotechnology. 6(4): 253-260.
- Moore NC. 2011. Silk moth's antenna inspires new nanotech tool with applications in Alzheimer's research. EurekAlert [Internet], Accessed 28-Feb-2011.
The silk-spinning organ of silkworms enhances crystallization and coagulation of the liquid silk product via shearing forces that help align the molecules.
"RAMSDEN (1938) observed that shearing of the viscous contents of silk glands between glass slides would cause it to coagulate. IIZUKA (1966) studied the effect of shear rate on the coagulation of fibroin solutions and also analyzed the dimensions of the spinneret of the silk worm Bombyx mori with regard to the shearing forces arising in this tube. He concluded that the shearing forces in the spinneret align molecules thus enhancing crystallization and also align the randomly oriented crystals thus promoting coagulation. The greater the shearing forces, the higher the degree of crystallinity of the product." (Wainwright 1976:77)
Learn more about this functional adaptation.
- Wainwright, S. A. 1982. Mechanical Design in Organisms. Princeton University Press.
Molecular Biology and Genetics
Barcode data: Bombyx mori
There are 59 barcode sequences available from BOLD and GenBank. Below is a sequence of the barcode region Cytochrome oxidase subunit 1 (COI or COX1) from a member of the species. See the BOLD taxonomy browser for more complete information about this specimen and other sequences.
-- end --
Download FASTA File
Statistics of barcoding coverage: Bombyx mori
Public Records: 51
Specimens with Barcodes: 64
Species With Barcodes: 1
Bombyx mori is currently not an endangered or threatened species; however, many animal rights activist groups object to their use in sericulture. One of the main things the activists are offended by is the silk industry's practice of boiling cocoons with living pupae inside in order to get the silk (envirolink 1997).
Relevance to Humans and Ecosystems
Economic Importance for Humans: Positive
In 1989, 74 thousand tons of silk were produced (Lepidoptera Part 2 1997). Even with each cocoon yeilding one half mile of fibers, that is an astounding amount of silk (The Animal World 1990). Bombyx mori is an incredibly important species to humans because we rely on their silk for our textile and clothing industries. For many years, China had a monopoly on the benifits of this industrious animal. In fact, Bombyx mori are one of the few animals that carried the death penalty as a punishment for smuggling them out of their native country (Lepidoptera Part 2 1997).
Bombyx mori are quite important animals in the science world as well. They are used in Australia for educational purposes in schools (Herbison-Evans 1997). Scientists in the field of sericulture are working on mapping their genes in hopes of improving the quality of the world's silk and expanding our knowledge of genetics in general. Bombyx mori were the animals in which pheromones were first discovered and named (Pines 1997).
|This article needs additional citations for verification. (July 2010)|
The silkworm is the larva or caterpillar of the domesticated silkmoth, Bombyx mori (Latin: "silkworm of the mulberry tree"). It is an economically important insect, being a primary producer of silk. A silkworm's preferred food is white mulberry leaves, but it may also eat the leaves of any other mulberry tree (i.e., Morus rubra or Morus nigra) as well as the Osage Orange. It is entirely dependent on humans for its reproduction and does not occur naturally in the wild. Sericulture, the practice of breeding silkworms for the production of raw silk, has been underway for at least 5,000 years in China, from where it spread to Korea and Japan, and later to India and the West. The silkworm was domesticated from the wild silkmoth Bombyx mandarina which has a range from northern India to northern China, Korea, Japan and far the eastern regions of Russia. The domesticated silkworm derives from Chinese rather than Japanese or Korean stock. It is unlikely that silkworms were domestically bred before the Neolithic age: it was not until then that the tools required to facilitate the manufacturing of larger quantities of silk thread had been developed. The domesticated B. mori and the wild B. mandarina can still breed and sometimes produce hybrids.:342
Mulberry silkworms can be categorized into 3 different, but connected groups or types. The major groups of silkworms fall under the univoltine ('uni-'=one, 'voltine'=brood frequency) and bivoltine categories. The Univoltine breed is generally linked with the geographical area within greater Europe. The eggs of this type hibernate during winter due to the cold climate, and cross fertilize only by spring, generating silk only once annually. The second type of breed is called Bivoltine and is normally found in Asian regions such as China, Japan, and Korea. The breeding process of this type takes place twice annually, a feat made possible through the slightly warmer climates and the resulting two lifecycles. The Polyvoltine breed of mulberry silkworm can only be located in the tropics. The eggs are laid by female moths and hatch within nine to twelve days, so the resulting type can have up to 8 separate life cycles throughout the year.
Eggs take about fourteen days to hatch into larvae, which eat continuously. They have a preference for white mulberry, having an attraction to the mulberry odorant cis-jasmone. They are not monophagous since they can eat other species of Morus as well as some other Moraceae. Their droppings are black. Hatchlings and second-instar larvae are called kego and chawki in India. They are covered with tiny black hairs. When the color of their heads turns darker, it indicates that they are about to molt. After molting, the instar phase of the silkworm emerges white, naked, and with little horns on the backs.
After they have molted four times, their bodies become slightly yellow and the skin becomes tighter. The larvae will then enter the pupa phase of their life cycle and enclose themselves in a cocoon made up of raw silk produced by the salivary glands. The cocoon provides a vital layer of protection during the vulnerable, almost motionless pupal state. Many other Lepidoptera produce cocoons, but only a few—the Bombycidae, in particular the Bombyx genus, and the Saturniidae, in particular the Antheraea genus—have been exploited for fabric production.
If the animal is allowed to survive after spinning its cocoon and through the pupa phase of its life cycle, it will release proteolytic enzymes to make a hole in the cocoon so that it can emerge as a moth. These enzymes are destructive to the silk and can cause the silk fibers to break down from over a mile in length to segments of random length, which seriously reduced the value of the silk threads. To prevent this, silkworm cocoons are boiled. The heat kills the silkworms and the water makes the cocoons easier to unravel. Often, the silkworm itself is eaten (see Cuisine).
As the process of harvesting the silk from the cocoon kills the larvae, sericulture has been criticized by animal welfare and rights activists. Mohandas Gandhi was critical of silk production based on the Ahimsa philosophy "not to hurt any living thing". This led to Gandhi's promotion of cotton spinning machines, an example of which can be seen at the Gandhi Institute. He also promoted Ahimsa silk, wild silk made from the cocoons of wild and semi-wild silk moths. Ahimsa silk is promoted in parts of Southern India for those who prefer not to wear silk produced by killing silkworms. Ahimsa silk is also known as peace silk. In the early 21st century the organization PETA has campaigned against silk.
The moth – the adult phase of the life cycle – cannot fly. Silkmoths have a wingspan of 3–5 cm (1.5–2 inches) and a white hairy body. Females are about two to three times bulkier than males (for they are carrying many eggs), but are similarly colored. Adult Bombycidae have reduced mouth parts and do not feed, though a human caretaker can also[clarification needed] feed them.
The cocoon is made of a thread of raw silk from 300 to about 900 meters (1,000 to 3,000 feet) long. The fibers are very fine and lustrous, about 10 micrometers (1/2,500th of an inch) in diameter. About 2,000 to 3,000 cocoons are required to make a pound of silk (0,4 kg). At least 70 million pounds of raw silk are produced each year, requiring nearly 10 billion pounds of cocoons. 
Due to its miniature size and ease of culture, the silkworm has become a model organism in the study of Lepidopteran and arthropod biology. Fundamental findings on pheromones, hormones, brain structures and physiology have been made with the silkworm. One example of this was the molecular identification of the first known pheromone, bombykol which required extracts from 500,000 individuals, due to the very small quantities of pheromone produced by any individual worm.
Currently, research is focusing on genetics of silkworms and the possibility of genetic engineering. Many hundreds of strains are maintained, and over 400 Mendelian mutations have been described. Another source suggests 1000 inbred domesticated strains are kept worldwide. One useful development for the silk industry are silkworms that can feed on food other than mulberry leaves, including an artificial diet. Research on the genome also raises the possibility of genetically engineering silkworms to produce proteins, including pharmacological drugs, in the place of silk proteins.
Kraig Biocraft Laboratories has used research from the Universities of Wyoming and Notre Dame in a collaborative effort to create a silkworm that is genetically altered to produce spider silk. In September 2010 it was announced at a press conference at the University of Notre Dame that the effort had been successful.
In January 2010, National University of Singapore together with Republic Polytechnic were in the process of creating "super" silk. The "super" silk are produced by coaxing "silkworms to spin stronger silk by exposing them (silkworms) to an electric field before they (silkworms) spin" . Silk produced through this new method is so strong as to be bullet-proof.
Researchers at Tufts developed scaffolds made of spongy silk that feel and look similar to human tissue. They are implanted during reconstructive surgery to support or restructure damaged ligaments, tendons, and other tissue. They also created implants made of silk and drug compounds which can be implanted under the skin for steady and gradual time release of medications. 
The domesticated variety, compared to the wild form, has increased cocoon size, growth rate and efficiency of its digestion. It has also gained tolerance to human presence and handling and living in crowded conditions. It also cannot fly and lacks fear of potential predators. These changes have made it entirely dependent upon humans for survival. It is kept in incubators (a machine in which a temperature can fixed for giving any organism heat like silkworms,new born babies,etc. are kept) so that the egg hatches.
Silkworm breeding 
The silkworm is one of the world's most genetically modified animals. Silkworms were first domesticated in China over 5000 years ago. Since then, the silk production capacity of the species has increased nearly tenfold. Silkworm is one of the few organisms wherein the principles of genetics and breeding were applied to harvest maximum output. It is next only to maize in exploiting the principles of heterosis and cross breeding. Silkworm breeding is aimed at the overall improvement of silkworm from a commercial point of view. The major objectives of silkworm breeding are improving fecundity (the egg laying capacity of a breed), healthiness of larvae, quantity of cocoon and silk production, disease resistance, etc. Healthiness of larvae leads to a healthy cocoon crop. Healthiness is dependent on factors such as better pupation rate, fewer dead larvae in the mountage, shorter larval duration (the shorter the larval duration, the lesser the chances of infection) and bluish tinged fifth instar larvae (it is observed that bluish colored fifth instar larvae are healthier than the reddish brown ones). Quantity of cocoon and silk produced is directly related to the pupation rate and larval weight. Healthier larvae have greater pupation rates and cocoon weights. Quality of cocoon and silk depends on a number of factors including genetics. Specific purposes apart from commercial purpose are given attention by advanced countries to breed development for specific purposes like sericin production, sex limited breeds, thin/thick filament production etc. Disease resistance breeding is important, as the major reason for crop losses is pathogen infection. Efforts are being made to select breeds which are tolerant or resistant to various pathogens. 
Silkworm raising as hobby and school science project 
In the USA, teachers may sometimes introduce insect life cycle to their students by raising silkworms in the classroom as a science project. Students have a chance to observe complete life cycles of insect from egg stage to larvae, pupa, moth and eggs.
The silkworm has been raised as a hobby in China and South Africa. Children often pass on the eggs, creating a noncommercial population. The experience provides children with the opportunity to witness the life cycle of silkworms. The practice of raising silkworms by children as pets has, in the non-silkfarming country of South Africa, led to the development of extremely hardy landraces of silkworm. This is because they are invariably subjected to hardships not encountered by commercially-farmed members of the species. However, these worms, not being selectively bred as such, are possibly inferior in silk production and may exhibit other undesirable traits.
The genome of the silkworm is mid-range with a genome size of ~432 Mb.
High genetic variability has been found in domestic lines of silkworms, though this is less than that among wild silkmoths (~83%). This suggests a single event of domestication, and that it happened over a short period of time, with a large number of wild worms having been collected for domestication. Major questions, however, remain unanswered: “Whether this event was in a single location or in a short period of time in several locations cannot be deciphered from the data”. Research also has yet to identify the area in China where domestication arose.
Like many insect species, silkworm pupae are eaten in some cultures (see Entomophagy), when it may go by the English phrase "ground cucumber".
- In Assam, India, they are boiled for extracting silk and the boiled pupa is taken directly with salt or fried with chilli pepper and or herbs to take as snack or dish.
- In Korea, they are boiled and seasoned to make a popular snack food known as beondegi.
- In China, street vendors sell roasted silkworm pupae.
- In Vietnam, this is known as con nhộng.
- Silkworms have also been proposed for cultivation by astronauts as space food on long-term missions.
Silkworm legends 
In China, there is a legend that the discovery of the silkworm's silk was by an ancient empress Lei Zu, the wife of the Yellow Emperor and the daughter of XiLing-Shi. She was drinking tea under a tree when a silk cocoon fell into her tea. As she picked it out and started to wrap the silk thread around her finger, she slowly felt a warm sensation. When the silk ran out, she saw a small larva. In an instant, she realized that this caterpillar larva was the source of the silk. She taught this to the people and it became widespread. There are many more legends about the silkworm.
The Chinese guarded their knowledge of silk, but, according to one story, a Chinese princess given in marriage to a Khotan prince brought to the oasis the secret of silk-manufacture, "hiding silkworms in her hair as part of her dowry", probably in the first half of the 1st century CE. It is also said that about AD 550, Christian monks smuggled silkworms, in a hollow stick, out of China and sold the secret to the Byzantine Empire. This is very unlikely, however, as there is no record that there were any Christians in China before the seventh century.
Silkworm diseases 
Nosema bombycis is a microsporidium that kills 100% of silkworms hatched from infected eggs. This disease can be carried over from worms to moths, then eggs and worms again. This microsporidium comes from the food that silkworms eat. If silkworms get this microsporidium in their worm stage, there are no visible symptoms. However, mother moths will pass the disease onto the eggs, and 100% of worms hatching from the diseased eggs will die in their worm stage. To prevent this disease, it is therefore extremely important to rule out all eggs from infected moths by checking the moth’s body fluid under a microscope.
Botrytis bassiana is a fungus that destroys the entire silkworm body. This fungus usually appears when silkworms are raised under cold conditions with high humidity. This disease is not passed on to the eggs from moths, as the infected silkworms cannot survive to the moth stage. This fungus can spread to other insects.
Grasserie: If grasserie is observed in chawkie stage, then the chawkie larvae must have been infected while hatching or during chawkie rearing. Infected eggs can be disinfected by cleaning their surface prior to hatching. Infections can occur as a result of improper hygiene in the chawkie rearing house. This disease develops faster in early instar rearing.
Pebrine is a disease caused by a parasitic microsporidian, Nosema bombycis Nageli. Diseased larvae show slow growth, an undersized, pale and flaccid body, and poor appetite. Tiny black spots appear on larval integument. Additionally, dead larvae will remain rubbery and do not undergo putrefaction after death.
Traditional Chinese medicine 
In Traditional Chinese medicine, silkworm is the source of the "stiff silkworm", which is made from dried 4th or 5th instar larvae which have died of white muscardine disease (a lethal fungal infection). It is believed to dispel flatulence, dissolve phlegm and relieve spasms.
See also 
- Grimaldi, David A.; Engel, Michael S. (2005). Evolution of the Insects. Cambridge University Press. ISBN 978-0-521-82149-0.
- Johnson, Sylvia (1989). Silkworms. Lerner Publications. ISBN 978-0-8225-9557-1. Children's book with lots of photos.
- Scoble, M. J. (1995). The Lepidoptera: Form, Function and Diversity. Princeton University Press. ISBN 978-0-19-854952-9.
- Yoshitake, N. (1968). "Phylogenetic aspects on the origin of Japanese race of the silkworm, Bombyx mori L.". Journal of Sericological Sciences of Japan 37: 83–87.
- Trevisan, Adrian. "Cocoon Silk: A Natural Silk Architecture". Sense of Nature.
- Wolchover, Natalie. "The Silk Renaissance". Seed Magazine.
- E. J. W. Barber (1992). Prehistoric Textiles: the Development of Cloth in the Neolithic and Bronze Ages with Special Reference to the Aegean. Princeton University Press. p. 31. ISBN 978-0-691-00224-8.
- K. P. Arunkumar, Muralidhar Metta & J. Nagaraju (2006). "Molecular phylogeny of silkmoths reveals the origin of domesticated silkmoth, Bombyx mori from Chinese Bombyx mandarina and paternal inheritance of Antheraea proylei mitochondrial DNA". Molecular Phylogenetics and Evolution 40 (2): 419–427. doi:10.1016/j.ympev.2006.02.023. PMID 16644243.
- Hideaki Maekawa, Naoko Takada, Kenichi Mikitani, Teru Ogura, Naoko Miyajima, Haruhiko Fujiwara, Masahiko Kobayashi & Osamu Ninaki (1988). "Nucleolus organizers in the wild silkworm Bombyx mandarina and the domesticated silkworm B. mori". Chromosoma 96 (4): 263–269. doi:10.1007/BF00286912.
- Brian K. Hall (2010). Evolution: Principles and Processes. Topics in Biology. Jones & Bartlett Learning. p. 400. ISBN 978-0-7637-6039-7.
- Trevisan, Adrian. "Cocoon Silk: A Natural Silk Architecture". Sense of Nature.
- "Mahatma Gandhi: 100 years", 1968, p. 349
- Silk Moths Fly Free Kusuma Rajaiah's Ahimsa project.
- Silk saree without killing a single silkworm Another article about Rajaiah and his methods.
- "Down and Silk: Birds and Insects Exploited for Fabric". PETA. Retrieved 06 January 2007.
- The International Silkworm Genome Consortium (2008). "The genome of a lepidopteran model insect, the silkworm Bombyx mori". Insect Biochemistry and Molecular Biology 38 (12): 1036–1045. doi:10.1016/j.ibmb.2008.11.004. PMID 19121390.
- "Kraig Biocraft Laboratories".
- "University of Notre Dame".
- "NUS: Creating "Super" Silk". R2m.nus.edu.sg. Retrieved 18 October 2011.
- "The Straits Times: Silk So Strong You Can Turn it into Bullet-proof vest" (PDF). Retrieved 18 October 2011.
- Marian R. Goldsmith, Toru Shimada & Hiroaki Abe (2005). "The genetics and genomics of the silkworm, Bombyx mori". Annual Review of Entomology 50: 71–100. doi:10.1146/annurev.ento.50.071803.130456. PMID 15355234.
- Hong-Song Yu1, Yi-Hong Shen, Gang-Xiang Yuan, Yong-Gang Hu1, Hong-En Xu1, Zhong-Huai Xiang and Ze Zhang. "Evidence of selection at melanin synthesis pathway loci during silkworm domestication". Molecular Biology and Evolution 28 (6): 1785–99 year=2011. doi:10.1093/molbev/msr002. PMID 21212153.
- Dennis Normile (2009). "Sequencing 40 Silkworm Genomes Unravels History of Cultivation". Science 325: 1058–1059. Bibcode:2009Sci...325.1058N. doi:10.1126/science.325_1058a. PMID 19713499.
- "Silkworm School Science Project Instruction" (PDF). Retrieved 18 October 2011.
- Kazuei Mita, Masahiro Kasahara, Shin Sasaki, Yukinobu Nagayasu, Tomoyuki Yamada, Hiroyuki Kanamori, Nobukazu Namiki, Masanari Kitagawa, Hidetoshi Yamashita, Yuji Yasukochi, Keiko Kadono-Okuda, Kimiko Yamamoto, Masahiro Ajimura, Gopalapillai Ravikumar, Michihiko Shimomura, Yoshiaki Nagamura, Tadasu Shin-i, Hiroaki Abe, Toru Shimada, Shinichi Morishita & Takuji Sasaki (2004). "The genome sequence of silkworm, Bombyx mori". DNA Research 11 (1): 27–35. doi:10.1093/dnares/11.1.27. PMID 15141943.
- Qingyou Xia, Yiran Guo, Ze Zhang, Dong Li, Zhaoling Xuan, Zhuo Li, Fangyin Dai, Yingrui Li, Daojun Cheng, Ruiqiang Li, Tingcai Cheng, Tao Jiang, Celine Becquet, Xun Xu, Chun Liu, Xingfu Zha, Wei Fan, Ying Lin, Yihong Shen, Lan Jiang, Jeffrey Jensen, Ines Hellmann, Si Tang, Ping Zhao, Hanfu Xu, Chang Yu, Guojie Zhang, Jun Li, Jianjun Cao, Shiping Liu, Ningjia He, Yan Zhou, Hui Liu, Jing Zhao, Chen Ye, Zhouhe Du, Guoqing Pan, Aichun Zhao, Haojing Shao, Wei Zeng, Ping Wu, Chunfeng Li, Minhui Pan, Jingjing Li, Xuyang Yin, Dawei Li, Juan Wang, Huisong Zheng, Wen Wang, Xiuqing Zhang, Songgang Li, Huanming Yang, Cheng Lu, Rasmus Nielsen, Zeyang Zhou, Jian Wang, Zhonghuai Xiang & Jun Wang (2009). "Complete resequencing of 40 genomes reveals domestication events and genes in silkworm (Bombyx)" (PDF). Science 326 (5951): 433–436. Bibcode:2009Sci...326..433X. doi:10.1126/science.1176620. PMID 19713493.
- Dennis Normile (2009). "Sequencing 40 silkworm genomes unravels history of cultivation". Science 325 (5944): 1058–1059. Bibcode:2009Sci...325.1058N. doi:10.1126/science.325_1058a. PMID 19713499.
- Choi, Charles Q. (13 January 2009). "Care for a Silkworm With Your Tang?". ScienceNOW Daily News. Retrieved 14 January 2009.
- Sarah Underhill Wisseman, Wendell S. Williams. Ancient Technologies and Archaeological Materials . Routledge, 1994. ISBN 2-88124-632-X. Page 131.
Further reading 
The Bombyx hybrid is a hybrid between a Bombyx mori female and a male Bombyx mandarina moth. They produce Silkworm larvae like all species of Bombyx. The larvae look a lot like the other variation, they are brown and the first half and gray at the bottom half, but they get larger black spots than the other variation, and they look like a normal Bombyx moth, but a bit darker. Instead, no hybrids are used for silk, (unlike a normal Silkworm of the Bombyx mori species), but for research. Bombyx mori females are much more likely to mate with a male Bombyx mandarina, but both species have to be kept in the same container. Since Bombyx Hybrids are much more common than the other variation more is known about them.
- "Evidence of Selection at Melanin Synthesis Pathway Loci during Silkworm Domestication". Mbe.oxfordjournals.org. 2011-01-06. http://mbe.oxfordjournals.org/content/28/6/1785.abstract. Retrieved 2011-10-18.
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