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BiologyInhabits deep parts of large rivers, with moderate to swift current (Ref. 59043). Adults live essentially in freshwater although some fish frequently occur in estuaries. Males are sexually mature between 9-15 years; females between 16-20 years (Ref. 11941). Spawns in main river channel over stone-gravel or gravel-sand bottom and with strong current (Ref. 59043).