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BiologyAnadromous and freshwater populations exist. At the sea, it occurs in shallow coastal and estuarine zones. In freshwaters, it inhabits deep parts of large rivers with moderate to swift current (Ref. 59043). Found mainly near the shore over sand and mud. Usually solitary, but swarms when hibernating or during spawning migrations. Occasionally forms schools (Ref. 3241). Feeds on benthic molluscs, crustaceans and small fishes. Spawns on stone or gravel bottom in large and deep rivers with strong current, 1-1.5 m/s (Ref. 59043). Mainly propagated through artificial reproduction.