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BiologyOccur in shallow coastal waters, and from brackish water to freshwater regions of rivers (Ref. 41299). Form schools (Ref. 40448); in larger aggregations during spawning, which takes place at sea (Ref. 3468). Feed on small algae, diatoms, forams, benthic polychaetes, crustaceans, mollusks, organic matter and detritus; fry feed on copepods and floating algae (Ref. 40448). Oviparous, eggs are pelagic and non-adhesive (Ref. 205). Caught mainly during the spawning season and is sold fresh; the roe is highly appreciated.