The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Afuang, L. E. (1994). Population ecology and distribution of Bufo marinus in the province of Isabela, Philippines. Unpublished Masters Thesis. Laguna: University of the Philippines at Los Banos.
  • Aguirre, W., S. Poss. 1999. "Indigenous Species in the Gulf of Mexico Ecosystem : Bufo marinus" (On-line). Accessed December 10, 1999 at
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  • Amphibian Species of the World V2.20 Database. The American Museum of Natural History. Department of Herpetology.">">
  • Andrew Snider and J. Bowler (1992) Longevity of Reptiles and Amphibians in North American Collections, Second Edition. Society for the Study of Amphibians and Reptiles .
  • Brown, G. P., Shine, R., Ward-Fear, G. (2010). ''Using a native predator (the meat ant, Iridomyrmex reburrus) to reduce the abundance of an invasive species (the cane toad, Bufo marinus) in tropical Australia.'' Journal of Applied Ecology, 47, 273â€"280.
  • Brown, R. M., & Diesmos A. C. (2002). Application of lineage-based species concepts to oceanic island frog populations: the effects of differing taxonomic philosophies on the estimation of Philippine biodiversity. The Silliman Journal. 42(1), 133-162.
  • Bruning, B., Phillips B. L., & Shine R. (2010). female mate choice in the Anura Turgid female toads give males the slip: a new mechanism of. Biology Letters.
  • Bruning, B., Phillips, B.L., Shine, R. (2010). ''Turgid female toads give males the slip: a new mechanism of female mate choice in the Anura.'' The Royal Society- Biology Letters
  • Bureau of Rural Sciences, C. 1998. "Commercial Use of Wild Animals: Cane Toad" (On-line). Accessed 19 June 2000 at
  • Cameron, E. 2002. "Cane Toads, Giant Toads or Marine Toads" (On-line). Australian Museum Online. Accessed July 28, 2005 at
  • Cogger, H. 1983. Reptiles and Amphibians of Australia. Sanibel, Florida: Ralph Curtis Press.
  • Crossland, M. R. (2000). ''Direct and indirect effects of the introduced toad Bufo marinus on populations of native anuran larvae in Australia.'' Ecography, 23(3), 283-290.
  • Crother, Brian I., Jeff Boundy, Frank T. Burbrink, et al. / Moriarty, John J., ed. 2008. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, With Comments Regarding Confidence in Our Understanding, Sixth Ed. Herpetological Circular No. 37
  • Crother, Brian I., Jeff Boundy, Frank T. Burbrink, et al. / Moriarty, John J., ed. 2008. Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, With Comments Regarding Confidence in Our Understanding, Sixth Ed. Herpetological Circular No. 37. 1-84
  • Davies, N. B., & Halliday T. R. (1977). Optimal mate selection in the toad Bufo bufo. Nature. 269, 56-58.
  • Drewes R. C., Roth B. (1981). ''Snail-eating frogs from the Ethiopian highlands: a new anuran specialization.'' Zoological Journal of the Linnean Society, 73, 267-287.
  • Easteal, S. (1963). ''Bufo marinus.'' Catalogue of American Amphibians and Reptiles. American Society of Ichthyologists and Herpetologists, 395.1-395.4.
  • Frost, D.R. 1985. AMPHIBIANS SPECIES OF THE WORLD: A Taxonomic and Geographical Reference. Published as a Joint Venture of Allen Press, Inc. and The Association Systematics Collections Lawrence, Kansas, U.S.A. 732 p..
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Gunther, A.C.L. 1900-02. Biologia Centrali-Americana. Reptilia and Batrachia. 1885-1902.
  • Holotype: Taylor, E. H. & Smith, H. M. 1945. Proc. U. S. Nat. Mus. 95 (3185): 553, figure 59.
  • Ingram, G. J., Covacevich J. (1990). ''Tropidonophis mairii vs. Bufo marinus. Memoirs of the Queensland Museum.'' , 29(396).
  • Invasive Species Specialist Group, 2005. "Bufo marinus" (On-line). Global Invasive Species Database. Accessed July 28, 2005 at
  • Krakauer, T. 1968. The Ecology of the Neotropical Toad, Bufo marinus in southern Florida. Herpetologica, 24(3): 214-221.
  • Lampo, M. and De Leo, G. A. (1998). ''The invasion ecology of the toad Bufo marinus: from South America to Australia.'' Ecological Applications, 8(2), 388-396.
  • Letnic, M., Webb J., & SHINE R. (2008). Invasive cane toads (Bufo marinus) cause mass mortality of freshwater crocodiles (Crocodylus johnstoni) in tropical Australia. Biological Conservation. 141, 1773–1782.
  • Llewelyn J., Phillips B. L., Shine R. (2009). ''Sublethal costs associated with the consumption of toxic prey by snakes.'' Austral Ecology, 34, 179â€"184.
  • Llewelyn, J., Schwarzkopf L., Alford R., Shine R. (2009). ''Something different for dinner? Responses of a native Australian predator (the keelback snake) to an invasive prey species (the cane toad).'' Biological Invasions
  • Maeda, N. and Matsui, M. (1990). Frogs and Toads of Japan, 2nd edition. Bun-Ichi Sogo Shuppan Co., Ltd., Tokyo, Japan.
  • Norman, D. 1998. Common Amphibians of Costa Rica/Anfibios Comunes de Costa Rica. U.S Fish & Wildlife Service, Asociación Conservacionista Yiski. 96 p.
  • Orrell T. (custodian) (2013). ITIS Global: The Integrated Taxonomic Information System (version Sept 12). In: Species 2000 & ITIS Catalogue of Life, 11th March 2013 (Roskov Y., Kunze T., Paglinawan L., Orrell T., Nicolson D., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Hernandez F., De Wever A., eds). Digital resource at Species 2000: Reading, UK.
  • Phillips, B.L., Brown, G.P., Shine, R. (2010). ''Evolutionarily accelerated invasions, the rate of dispersal evolves upwards during the range advance of cane toads.'' Journal of Evolutionary Biology
  • Prohl, H. 1997. Los Anfibios de Hitoy Cerere, Costa Rica. Proyecto Namasol, Cooperación Técnica Bilateral Holanda-Costa Rica. San José, Costa Rica. 66 p.
  • Savage, J.M. 1973. A Preliminary Handlist of the Herpetofauna of Costa Rica. s.d.. 17 p..
  • Savage, J.M. 2002. The Amphibians and Reptiles of Costa Rica. The University of Chicago Press. China. 934 pp.
  • Savage, J.M. y J. Villa. 1986. Introduction to the Herpetofauna of Costa Rica/Introducción a la Herpetofauna de Costa Rica. Society for the Studies of Amphibians and Reptiles, Athens, 207.
  • Seebacher, F. and Alford, R. A. (1999). ''Movement and microhabitat use of a terrestrial amphibian (Bufo marinus) on a tropical island: seasonal variation and environmental correlates.'' Journal of Herpetology, 33(2), 208-214.
  • Shine, R. (2006). ''The Ecological Impact of Invasive Cane Toads (Bufo marinus) in Australia'' Quarterly Review of Biology: in press.
  • Taylor, E.H. 1952. A review of frogs and toads of Costa Rica. Uni. Kansas Sci. Bull. 35: 577-942.
  • Universidad de Costa Rica. s.f.. CATALOGO DE HERPETOLOGIA. Museo de Zoología, Escuela de Biología, Universidad de Costa Rica. s.d..
  • Williams, C. R., Brodie J.. E. D., Tyler M. J., & Walker S. J. (2000). Antipredator Mechanisms of Australian Frogs. Journal of Herpetology. 34, 431-443.
  • Zug, G.R. y P.B. Zug. 1979. The marine toad, Bufo marinus: A natural history resumé of native population. Smithsonian Contrib. Zool. 284: 1-58.
  • Zug, N. 1991. Bufo marinus (Sapo Grande, Sapo, Giant Toad, Marine Toad). Pp. 390-392. In: D.H. Janzen (ed.). Historia Natural de Costa Rica. Editorial de la Universidad de Costa Rica. San José, Costa Rica.


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