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Range DescriptionThis species occurs in northeastern and northern Madagascar, with records from Marojejy at 300 m asl and Est Betaidambo in the Analabe Rainforest along Ramena River (west of Tsaratanana Reserve National Integral) at 700 m asl. It was described in 1999, based on a single specimen in the collections of the Zoological Institute of the University of Antananarivo. The original description was based on a colour slide, but the holotype has since been found and redescriptions have been published (Vences et al. 2004).