The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 2004. database, Amphibian Species of the World: an Online Reference
  • Boettger, O. (1892): Katalog der Batrachier-Sammlung im Museum der Senckenbergischen Naturforschenden Gesellschaft in Frankfurt am Main.
  • Coloma, L. A., Lotters, S. A., and Salas, A. W. (2000). ''Taxonomy of the Atelopus ignescens complex (Anura: Bufonidae): Designation of a neotype of Atelopus ignescens and recognition of Atelopus exiguus.'' Herpetologica, 56(3), 303-324.
  • Coloma, L. A., Lötters, S., Duellman, W.E., and Miranda-Leiva, A. (2007). ''A taxonomic revision of Atelopus pachydermus, and description of two new (extinct?) species of Atelopus from Ecuador (Anura: Bufonidae).'' Zootaxa, 1557, 1-32.
  • Coloma, L. A., and Ron, S. R. (2001). Ecuador Megadiverso: Anfibios, Reptiles, Aves, Mamiferos / Megadiverse Ecuador: Amphibians, Reptiles, Birds, And Mammals. Serie de divulgación del Centro de Biodiversidad y Ambiente. Quito, Ecuador.
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Gray, P. (1983). Morphometrics of the Atelopus ignescens Complex (Anura: Bufonidae). M. S. Thesis. University of Kansas, Lawrence, Kansas.
  • Guayasamin, J. M., Bonaccorso, E., Duellman, W. E., and Coloma, L. A. (2010). ''Genetic differentiation in the nearly extinct harlequin frogs (Bufonidae: Atelopus), with emphasis on the Andean Atelopus ignescens and A. bomolochos species complexes.'' Zootaxa , 2574, 55-68.
  • La Marca, E., Lötters, S., Puschendorf, R., Ibáñez, R., Rueda-Almonacid, J. V., Schulte, R., Marty, C., Castro, F., Manzanilla-Puppo, J., García-Pérez, J. E., Bolaños, F., Chaves, G., Pounds, J. A., Toral, E., and Young, B. E. (2005). ''Catastrophic population declines and extinctions in neotropical harlequin frogs (Bufonidae: Atelopus).'' Biotropica, 37(2), 190-201.
  • Moore, R. (2007). ''In search of Atelopus exiguus.'' Froglog, 83, 1-2.
  • Orrell T. (custodian) (2013). ITIS Global: The Integrated Taxonomic Information System (version Sept 12). In: Species 2000 & ITIS Catalogue of Life, 11th March 2013 (Roskov Y., Kunze T., Paglinawan L., Orrell T., Nicolson D., Culham A., Bailly N., Kirk P., Bourgoin T., Baillargeon G., Hernandez F., De Wever A., eds). Digital resource at Species 2000: Reading, UK.
  • Read, M. (1986). ''Reptiles and amphibians.'' Río Mazán Project, 1986 Report. F. Robinson, eds., Department of Plant Sciences, Oxford University, Oxford, U. K..
  • Ron, S.R., and Merino-Viteri, A. (2000). ''Amphibian declines in Ecuador: overview and first report of chytridiomycosis from South America.'' Froglog, 42, 2-3.
  • Stuart, S., Hoffmann, M., Chanson, J., Cox, N., Berridge, R., Ramani, P., and Young, B. (eds) (2008). Threatened Amphibians of the World. Lynx Edicions, IUCN, and Conservation International, Barcelona, Spain; Gland, Switzerland; and Arlington, Virginia, USA.
  • Toral, E., and and Frenkel, C. (2006). ''Population state of Atelopus exiguus in the South Ecuadorian Andes (Abstracts).'' Joint Meeting of Ichthyologists & Herpetologists, 2006. M. A. Donnelly and K. E. Cooper, eds., New Orleans, LA, 111-112.


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