The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • Amphibian Species of the World V2.20 Database. The American Museum of Natural History. Department of Herpetology.">">
  • Cannatella, D.C. and Duellman, W.E. (1984). ''Leptodactylid frogs of the Physalaemus pustulosus group.'' Copeia, 1984(4), 902-921.
  • Costa Rica, Asamblea Legislativa. Decreto No. 26435-MINAE. La Gaceta, Diario Oficial (3 de Diciembre 1997).
  • Costa Rica, Asamblea Legislativa. Ley Orgánica del Ambiente, No. 7554. La Gaceta, Diario Oficial, No. 215 (13 de Noviembre 1995).
  • Costa Rica, Ministerio de Ambiente y Energía (MINAE). 1997. Ley de Conservación de la Vida Silvestre y su Reglamento, No. 7317. 2 da. Edición. EDITEC EDITORES, S.A.. San José, Costa Rica. 134 p.
  • Dalgetty L; Kennedy MW. 2010. Building a home from foam--túngara frog foam nest architecture and three-phase construction process. Biology Letters. 6(3): 293-6.
  • Downer, J. 2002. Weird Nature: An Astonishing Exploration of Nature's Strangest Behavior. Ontario: Firefly Books.
  • Fleming RI; Mackenzie CD; Cooper A; Kennedy MW. 2009. Foam nest components of the túngara frog: a cocktail of proteins conferring physical and biological resilience. Proc. Biol. Sci. 276(1663): 1787-95.
  • Frost, D.R. 1985. AMPHIBIANS SPECIES OF THE WORLD: A Taxonomic and Geographical Reference. Published as a Joint Venture of Allen Press, Inc. and The Association Systematics Collections Lawrence, Kansas, U.S.A. 732 p..
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Gunther, A.C.L. 1900-02. Biologia Centrali-Americana. Reptilia and Batrachia. 1885-1902.
  • Lahanas, P.N.. 1994. The Function of Near Neighbors in Decreasing Call Latency Period by the Túngara Frog, Physalaemus pustulosus. Biotropica 27(2): 262-265.
  • Marsh, D.M., Fegraus, E.H., and Harrison, S. (1999). ''Effects of breeding pond isolation on the spatial and temporal dynamics of pond use by the Tungara Frog, Physalaemus pustulosus.'' Journal of Animal Ecology, 68, 804-814.
  • Morelle R. 2010. How to whip up the perfect frothy frog 'meringue' nest. BBC News [Internet],
  • Norman, D. 1998. Common Amphibians of Costa Rica/Anfibios Comunes de Costa Rica. U.S Fish & Wildlife Service, Asociación Conservacionista Yiski. 96 p.
  • Orrell T. (custodian) (2015). ITIS Global: The Integrated Taxonomic Information System (version Mar 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Rand, A.S. 1991. Physalaemus pustulosus (Rana, Sapito Túngara, Foam Toad, Mud-puddle Frog). Pp. 417-418, In: D.H. Janzen (ed.). Historia Natural de Costa Rica. Editorial de la Universidad de Costa Rica. San José, Costa Rica.
  • Ryan, M.J. (1985). The Túngara Frog: A Study in Sexual Selection and Communication. University of Chicago Press, Chicago.
  • Ryan, M.J., Rand, A.S., and Weigt, L.A. (1996). ''Allozyme and advertisement call variation in the Tungara frog, Physalaemus pustulosus.'' Evolution, 50(6), 2435-2453.
  • Santos-Barrera, G., Solís, F., Ibáñez, R., Wilson, L. D., Savage, J., Bolaños, F., Lee, J., Chaves, G., Señaris, C., Acosta-Galvis, A., and Hardy, J. 2008. Engystomops pustulosus. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.1. Downloaded on 29 March 2010.
  • Savage, J.M. 1973. A Preliminary Handlist of the Herpetofauna of Costa Rica. s.d.. 17 p..
  • Savage, J.M. 2002. The amphibians and reptiles of Costa Rica. The University of Chicago Press. China. 934 p.
  • Savage, J.M. y J. Villa. 1986. Introduction to the Herpetofauna of Costa Rica/Introducción a la Herpetofauna de Costa Rica. Society for the Studies of Amphibians and Reptiles, Athens, 207.
  • Taylor, E.H. 1952. A review of frogs and toads of Costa Rica. Uni. Kansas Sci. Bull. 35: 577-942.
  • Tárano, Z. (1998). ''Cover and ambient light influence nesting preferences in the Túngara Frog Physalaemus pustulosus.'' Copeia, 1998(1), 250-251.
  • Universidad de Costa Rica. s.f.. CATALOGO DE HERPETOLOGIA. Museo de Zoología, Escuela de Biología, Universidad de Costa Rica. s.d..
  • Wendell, D., Todd, J., and Montemagno, C. (2010). ''Artificial photosynthesis in ranaspumin-2 based foam.'' Nano Letters, DOI: 10.1021/nl100550k.


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