The following bibliography has been generated by bringing together all references provided by our content partners. There may be duplication.


  • 1996. database, NODC Taxonomic Code
  • 2004. database, Amphibian Species of the World: an Online Reference
  • Amphibian Species of the World V2.20 Database. The American Museum of Natural History. Department of Herpetology.">">
  • Berger, L., Speare, R., Daszak, P., Green, D. E., Cunningham, A. A., Goggin, C. L., Slocombe, R., Ragan, M. A., Hyatt, A. D., McDonald, K. R., Hines, H. B., Lips, K. R., Marantelli, G., and Parkes, H. (1998). "Chytridiomycosis causes amphibian mortality associated with population declines in the rain forests of Australia and Central America." Proceedings of the National Academy of Sciences of the United States of America, 95(15), 9031-9036.
  • Capula, M. 1989. Guide to Reptiles and Amphibians of the World. New York: Simon and Schuster.
  • Cope, E.D.. 1875. On the Batrachia and Reptilia of Costa Rica. J. Acad. Nat. Sci. Phila.,. Ser. 2, (8): 93-157 (Reimpreso por O'Bios).
  • Costa Rica, Asamblea Legislativa. Decreto No. 26435-MINAE. La Gaceta, Diario Oficial (3 de Diciembre 1997).
  • Costa Rica, Asamblea Legislativa. Ley Orgánica del Ambiente, No. 7554. La Gaceta, Diario Oficial, No. 215 (13 de Noviembre 1995).
  • Costa Rica, Ministerio de Ambiente y Energía (MINAE). 1997. Ley de Conservación de la Vida Silvestre y su Reglamento, No. 7317. 2 da. Edición. EDITEC EDITORES, S.A.. San José, Costa Rica. 134 p.
  • Crump, M. 1988. Aggresion in harlequin frogs: male-male competition and a possible conflict of interest between the sexes. Anim. Behav. 36: 1064-1077.
  • Crump, M. L., and Pounds, J. A. (1985). ''Lethal parasitism of an aposematic anuran (Atelopus varius) by Notochaeta bufonivora (Diptera: Sarcophagidae).'' Journal of Parasitology, 71(5), 588-591.
  • Crump, M. y J.A. Pounds. 1985. Lethal parasitism of an aposematic anuran (Atelopus varius) by Notohaeta bufonivora (Diptera: Sarcophagidae). J. Parasitol. 71: 588-591.
  • Crump, M. y J.A. Pounds. 1989. Temporal variation in the dispersion of a tropical anuran. Copeia 1989 (1): 299-211.
  • Crump, M.. 1986. Homing and site fidelity in a Neotropical frog, Atelopus varius (Bufonidae). Copeia 1986(2): 438-444.
  • Crump, M.L. (1988). ''Aggression in Harlequin Frogs: male-male competition and a possible conflict of interest between the sexes.'' Animal Behaviour, 36(4), 1064-1077.
  • Daly, J.W., Padgett, W.L., Saunders, R.L., and Cover, J.F., Jr. (1997). ''Absence of tetrodotoxins in a captive-raised riparian frog, Atelopus varius.'' Toxicon, 35(5), 705-709.
  • Frank & Kate's Web Page (Online source) External link.
  • Frost, D.R. 1985. AMPHIBIANS SPECIES OF THE WORLD: A Taxonomic and Geographical Reference. Published as a Joint Venture of Allen Press, Inc. and The Association Systematics Collections Lawrence, Kansas, U.S.A. 732 p..
  • Frost, Darrel R. 1997. Amphibian Species of the World, A Taxonomic and Geographic Reference. Second edition
  • Frost, Darrel R. 1997. Amphibian Species of the World, A Taxonomic and Geographic Reference. Second edition
  • Frost, Darrel R. 2009. database. Amphibian Species of the World: an Online Reference v5.3
  • Gunther, A.C.L. 1900-02. Biologia Centrali-Americana. Reptilia and Batrachia. 1885-1902.
  • Hayes, M., J. Pounds, W. Timmerman. undated (Herp. Circ. No. 17). An Annotated List and Guide to the Amphibians and Reptiles of Monteverde, Costa Rica. Tyler, Texas: Soc. for the Study of Amphibians and Reptiles.
  • Holdridge, L. R. (1967). Life Zone Ecology. Tropical Science Center, San Jose, Costa Rica.
  • Kim, Y., G. B. Brown y F. Fuhrman. 1975. Tetrodotoxin: Ocurrence in Atelopid Frogs of Costa Rica. Science 189(4197): 151-152.
  • Levell, J. 1997. A Field Guide to Reptiles and the Law. Lanesboro, Minnesota: Serpent's Tale.
  • Orrell T. (custodian) (2015). ITIS Global: The Integrated Taxonomic Information System (version Mar 2015). In: Species 2000 & ITIS Catalogue of Life, 26th August 2015 (Roskov Y., Abucay L., Orrell T., Nicolson D., Kunze T., Flann C., Bailly N., Kirk P., Bourgoin T., DeWalt R.E., Decock W., De Wever A., eds). Digital resource at Species 2000: Naturalis, Leiden, the Netherlands. ISSN 2405-8858.
  • Pounds A. y M. L. Crump. 1987. Harlequin Frogs Along a Tropical Montane Stream: Aggregation and the Risk of Predation by Frog-Eating Flies. Biotropica 19(4): 306-309.
  • Pounds, A. y M.L. Crump. 1994. Amphibians declines and climate disturbance: the case of the golden toad and the harlequín frog. Cons. Bio. 8: 72-85.
  • Pounds, J. A., Fogden, M. P. L., and Campbell, J. H. (1999). ''Biological response to climate change on a tropical mountain.'' Nature, 398(6728), 611-615.
  • Pounds, J. A., and Crump, M. L. (1987). ''Harlequin Frogs along a tropical montane stream: Aggregation and the risk of predation by frog-eating flies.'' Biotropica, 19(4), 306-309.
  • Pounds, J. A., and Crump, M. L. (1989). ''Temporal variation in the dispersion of a tropical anuran.'' Copeia, 1989(1), 209-211.
  • Pounds, J. A., and Crump, M. L. (1994). ''Amphibian declines and climate disturbance: The case of the Golden Toad and the Harlequin Frog.'' Conservation Biology, 8(1), 72-85.
  • Rodríguez, J. y J. Hernández. 1999. Especies de Flora y Fauna Silvestre con Poblaciones Reducidas y en Peligro de Extinción. Sistema Nacional de Areas de Conservación, Ministerio de Ambiente y Energía (MINAE). San José, Costa Rica. 12 p.
  • Savage, J.M. (1972). ''The harlequin frogs, genus Atelopus, of Costa Rica and western Panama.'' Herpetologica, 28(2), 77-94.
  • Savage, J.M. 1972. The harlequín frogs, genus Atelopus of Costa Rica and Western Panamá. Herpetologica 28(2): 77-94.
  • Savage, J.M. 1973. A Preliminary Handlist of the Herpetofauna of Costa Rica. s.d.. 17 p..
  • Savage, J.M. y J. Villa. 1986. Introduction to the Herpetofauna of Costa Rica/Introducción a la Herpetofauna de Costa Rica. Society for the Studies of Amphibians and Reptiles, Athens, 207.
  • Staniszewski, M. 1995. Amphibians in Captivity. Neptune, New Jersey: T.F.H. Publ., Inc..
  • Starret, P. 1967. Observations on the life history of frogs of the family Atelopodidae. Herpetologica 23(3): 195-204.
  • Taylor, E. H.. 1955. Additions to the Known Herpetological Fauna of Costa Rica With Comments on Other Species No. ll. The University of Kansas Science Bulletin XXXVll: 499-575.
  • Taylor, E.H. 1952. A review of frogs and toads of Costa Rica. Uni. Kansas Sci. Bull. 35: 577-942.
  • U.S. Geological Survey, 1999. USGS Issues Wildlife Health Alert: Chytrid Fungus Infection Associated With Deaths of Threatened Boreal Toads in Colorado. Froglog: Newsletter of the Declining Amphibian Populations Task Force, Dec. 1999: No. 36: 1-2.
  • Universidad de Costa Rica. s.f.. CATALOGO DE HERPETOLOGIA. Museo de Zoología, Escuela de Biología, Universidad de Costa Rica. s.d..


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