Overview

Comprehensive Description

Description

Trees or shrubs. Flowers appearing before or after the leaves, in usually erect catkins, bracts entire or toothed; each flower with 1-2 fleshy glands; stamens 2-many.
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© Mark Hyde, Bart Wursten and Petra Ballings

Source: Flora of Zimbabwe

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Molecular Biology and Genetics

Molecular Biology

Statistics of barcoding coverage

Barcode of Life Data Systems (BOLD) Stats
                                        
Specimen Records:2,298Public Records:1,256
Specimens with Sequences:1,990Public Species:214
Specimens with Barcodes:1,948Public BINs:0
Species:381         
Species With Barcodes:340         
          
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© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au035

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au034

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au032

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au011

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au010

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au002

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au001

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au095

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au094

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au093

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au089

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au088

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au087

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au082

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au081

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au080

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au079

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au078

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au077

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
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© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au076

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au075

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au074

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
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© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au047

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au046

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au045

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au043

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au036

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au031

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au018

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au014

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au013

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au004

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au003

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au092

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au091

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au090

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au086

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au085

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au069

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au068

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au064

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
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© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au063

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
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© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix sp.

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 20
Specimens with Barcodes: 31
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au056

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au071

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au070

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix sp. 5

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
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© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix sp. 4

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
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© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix sp. 3

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix sp. 2

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au058

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au053

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au052

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au051

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au050

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au048

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au030

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au029

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au020

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au062

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Default rating: 2.5 of 5

Statistics of barcoding coverage: Salix Au060

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au059

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Statistics of barcoding coverage: Salix Au023

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Default rating: 2.5 of 5

Statistics of barcoding coverage: Salix Au022

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Default rating: 2.5 of 5

Statistics of barcoding coverage: Salix sp. 1

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Default rating: 2.5 of 5

Statistics of barcoding coverage: Salix Au057

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Default rating: 2.5 of 5

Statistics of barcoding coverage: Salix Au055

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 2
Species With Barcodes: 1
Creative Commons Attribution 3.0 (CC BY 3.0)

© Barcode of Life Data Systems

Source: Barcode of Life Data Systems (BOLD)

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Default rating: 2.5 of 5

Statistics of barcoding coverage: Salix Au017

Barcode of Life Data Systems (BOLDS) Stats
Public Records: 0
Specimens with Barcodes: 1
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Collection Sites: world map showing specimen collection locations for Salix

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Wikipedia

Willow

For other uses, see Willow (disambiguation).
"Willow tree" redirects here. For Willow tree figurines, see Willow Tree (figurines).
"Salix" redirects here. For other uses, see Salix (disambiguation).

Willows, sallows, and osiers form the genus Salix, around 400 species[2] of deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions of the Northern Hemisphere. Most species are known as willow, but some narrow-leaved shrub species are called osier, and some broader-leaved species are referred to as sallow (from Old English sealh, related to the Latin word salix, willow). Some willows (particularly arctic and alpine species) are low-growing or creeping shrubs; for example, the dwarf willow (Salix herbacea) rarely exceeds 6 cm (2 in) in height, though it spreads widely across the ground.

Description[edit]

At the base of the petiole a pair of stipules form. These may fall in spring, or last for much of the summer or even for more than one year (marcescence).

Willows all have abundant watery bark sap, which is heavily charged with salicylic acid, soft, usually pliant, tough wood, slender branches, and large, fibrous, often stoloniferous roots. The roots are remarkable for their toughness, size, and tenacity to life, and roots readily grow from aerial parts of the plant.

The leaves are typically elongated, but may also be round to oval, frequently with serrated edges. Most species are deciduous; semievergreen willows; coriaceous leaves are rare, e.g. Salix micans and S. australior in the eastern Mediterranean. All the buds are lateral; no absolutely terminal bud is ever formed. The buds are covered by a single scale. Usually, the bud scale is fused into a cap-like shape, but in some species it wraps around and the edges overlap.[3] The leaves are simple, feather-veined, and typically linear-lanceolate. Usually they are serrate, rounded at base, acute or acuminate. The leaf petioles are short, the stipules often very conspicuous, resembling tiny, round leaves, and sometimes remaining for half the summer. On some species, however, they are small, inconspicuous, and caducous (soon falling). In color, the leaves show a great variety of greens, ranging from yellowish to bluish.

Flowers[edit]

Young male catkin

Willows are dioecious, with male and female flowers appearing as catkins on different plants; the catkins are produced early in the spring, often before the leaves, or as the new leaves open.

The staminate (male) flowers are without either calyx or corolla; they consist simply of stamens, varying in number from two to 10, accompanied by a nectariferous gland and inserted on the base of a scale which is itself borne on the rachis of a drooping raceme called a catkin, or ament. This scale is square, entire, and very hairy. The anthers are rose-colored in the bud, but orange or purple after the flower opens; they are two-celled and the cells open longitudinally. The filaments are threadlike, usually pale brown, and often bald.

The pistillate (female) flowers are also without calyx or corolla, and consist of a single ovary accompanied by a small, flat nectar gland and inserted on the base of a scale which is likewise borne on the rachis of a catkin. The ovary is one-celled, the style two-lobed, and the ovules numerous.

Cultivation[edit]

Almost all willows take root very readily from cuttings or where broken branches lie on the ground. The few exceptions include the goat willow (Salix caprea) and peachleaf willow (Salix amygdaloides). One famous example of such growth from cuttings involves the poet Alexander Pope, who begged a twig from a parcel tied with twigs sent from Spain to Lady Suffolk. This twig was planted and thrived, and legend has it that all of England's weeping willows are descended from this first one.[4][5]

Willows are often planted on the borders of streams so their interlacing roots may protect the bank against the action of the water. Frequently, the roots are much larger than the stem which grows from them.

Hybrids[edit]

Willows are very cross-fertile, and numerous hybrids occur, both naturally and in cultivation. A well-known ornamental example is the weeping willow (Salix × sepulcralis), which is a hybrid of Peking willow (Salix babylonica) from China and white willow (Salix alba) from Europe.

The hybrid cultivar 'Boydii' has gained the Royal Horticultural Society's Award of Garden Merit.[6]

Ecological issues[edit]

Knotted willow and woodpile in the Bourgoyen-Ossemeersen, Ghent, Belgium

Willows are used as food plants by the larvae of some Lepidoptera species, such as the Mourning Cloak butterfly.[7] Ants, such as wood ants, are common on willows inhabited by aphids, coming to collect aphid honeydew, as sometimes do wasps.

A small number of willow species were widely planted in Australia, notably as erosion-control measures along watercourses. They are now regarded as invasive weeds, and many catchment management authorities are removing and replacing them with native trees.[8][9]

Willow roots spread widely and are very aggressive in seeking out moisture; for this reason, they can become problematic when planted in residential areas, where the roots are notorious for clogging French drains, drainage systems, weeping tiles, septic systems, storm drains, and sewer systems, particularly older, tile, concrete, or ceramic pipes. Newer, PVC sewer pipes are much less leaky at the joints, and are therefore less susceptible to problems from willow roots; the same is true of water supply piping.[10][11]

Pests and diseases[edit]

Willow species are hosts to more than a hundred aphid species, belonging to Chaitophorus and other genera,[12] forming large colonies to feed on plant juices, on the underside of leaves in particular.[13] Rust, caused by fungi of genus Melampsora, is known to damage leaves of willows, covering them with orange spots.[14]

Uses[edit]

Medicine[edit]

The leaves and bark of the willow tree have been mentioned in ancient texts from Assyria, Sumer and Egypt[15] as a remedy for aches and fever,[16] and in Ancient Greece the physician Hippocrates wrote about its medicinal properties in the fifth century BC. Native Americans across the Americas relied on it as a staple of their medical treatments. It provides temporary pain relief. Salicin is metabolized into salicylic acid in the human body, and is a precursor of aspirin.[17] In 1763, its medicinal properties were observed by the Reverend Edward Stone in England. He notified the Royal Society, which published his findings. The active extract of the bark, called salicin, was isolated to its crystalline form in 1828 by Henri Leroux, a French pharmacist, and Raffaele Piria, an Italian chemist, who then succeeded in separating out the compound in its pure state. In 1897, Felix Hoffmann created a synthetically altered version of salicin (in his case derived from the Spiraea plant), which caused less digestive upset than pure salicylic acid. The new drug, formally acetylsalicylic acid, was named Aspirin by Hoffmann's employer Bayer AG. This gave rise to the hugely important class of drugs known as nonsteroidal anti-inflammatory drugs (NSAIDs).

Willow has been listed as one of the 38 substances used to prepare Bach flower remedies,[18] a kind of alternative medicine promoted for its effect on health. However according to Cancer Research UK, "there is no scientific evidence to prove that flower remedies can control, cure or prevent any type of disease, including cancer".[19]

Manufacturing[edit]

Some of humans' earliest manufactured items may have been made from willow. A fishing net made from willow dates back to 8300 BC.[20] Basic crafts, such as baskets, fish traps, wattle fences and wattle and daub house walls, were often woven from osiers (rod-like willow shoots, often grown in coppices). One of the forms of Welsh coracle traditionally uses willow in the 'lats'. Thin or split willow rods can be woven into wicker, which also has a long history. The relatively pliable willow is less likely to split while being woven than many other woods, and can be bent around sharp corners in basketry. Willow wood is also used in the manufacture of boxes, brooms, cricket bats (grown from certain strains of white willow), cradle boards, chairs and other furniture, dolls, flutes, poles, sweat lodges, toys, turnery, tool handles, veneer, wands and whistles. In addition, tannin, fibre, paper, rope and string can be produced from the wood.

Other[edit]

Male catkin of Salix cinerea with bee
Willow tree with woodbine honeysuckle
The willow tree as seen as the main part of an heraldic escutcheon over the main portal of a patrician house belonging to the Salis family in Chur, Switzerland, circa 1750
Environmental art installation "Sandworm" in the Wenduine Dunes, Belgium, made entirely out of willow
  • Agriculture: Willows produce a modest amount of nectar from which bees can make honey, and are especially valued as a source of early pollen for bees. Poor people at one time often ate willow catkins that had been cooked to form a mash.[21]
  • Art: Willow is used to make charcoal (for drawing) and in living sculptures. Living sculptures are created from live willow rods planted in the ground and woven into shapes such as domes and tunnels. Willow stems are used to weave baskets and three-dimensional sculptures, such as animals and figures. Willow stems are also used to create garden features, such as decorative panels and obelisks.

Culture[edit]

  • Culture: In China, some people carry willow branches with them on the day of their Tomb Sweeping or Qingming Festival. Willow branches are also put up on gates and/or front doors, which they believe help ward off the evil spirits that wander on Qingming. Legend states that on Qingming Festival, the ruler of the underworld allows the spirits of the dead to return to earth. Since their presence may not always be welcome, willow branches keep them away.[26] In traditional pictures of the Goddess of Mercy Guanyin, she is often shown seated on a rock with a willow branch in a vase of water at her side. The Goddess employs this mysterious water and the branch for putting demons to flight. Taoist witches also use a small carving made from willow wood for communicating with the spirits of the dead. The image is sent to the nether world, where the disembodied spirit is deemed to enter it, and give the desired information to surviving relatives on its return.[27] The willow is a famous subject in many East Asian nations' cultures, particularly in pen and ink paintings from China and Japan.
A gisaeng (Korean geisha) named Hongrang, who lived in the middle of the Joseon Dynasty, wrote the poem "By the willow in the rain in the evening", which she gave to her parting lover (Choi Gyeong-chang).[28] Hongrang wrote:

"...I will be the willow on your bedside."

In Japanese tradition, the willow is associated with ghosts. It is popularly supposed that a ghost will appear where a willow grows. Willow trees are also quite prevalent in folklore and myths.[29][30]
In English folklore, a willow tree is believed to be quite sinister, capable of uprooting itself and stalking travellers. The Viminal Hill, one of the Seven Hills Of Rome, derives it name from the Latin word for osier, viminia (pl.).
  • Hans Christian Andersen wrote a story called "Under the Willow Tree" (1853) in which children ask questions of a tree they call "willow-father", paired with another entity called "elder-mother".[31]
  • Old Man Willow in J. R. R. Tolkien's legendarium, appearing in The Lord of the Rings.
  • "Green Willow" is a Japanese ghost story in which a young samurai falls in love with a woman called Green Willow who has a close spiritual connection with a willow tree.[32] "The Willow Wife" is another, not dissimilar tale.[33] "Wisdom of the Willow Tree" is an Osage Nation story in which a young man seeks answers from a willow tree, addressing the tree in conversation as 'Grandfather'.[34]
  • In "The Secret of Salix Babylonicus," set in the Babylonian exile, the willow is portrayed benignly as a symbol of sympathy in grief and perseverance in suffering, an upright character who shows solidarity with the exiles and comes to express the spirit of the psalms.

Selected species[edit]

Main article: List of Salix species

The genus Salix is made up of around 400 species[2] of deciduous trees and shrubs:

See also[edit]

References[edit]

  1. ^ "Genus Salix (willows)". Taxonomy. UniProt. Retrieved 2010-02-04. 
  2. ^ a b c Mabberley, D.J. 1997. The Plant Book, Cambridge University Press #2: Cambridge.
  3. ^ George W. Argus. The Genus Salix (Salicaceae) in the Southeastern United States. Systematic Botany Monographs 9. 
  4. ^ Leland, John (2005) Aliens in the Backyard: Plant and Animal Imports into America, p. 70. Univ of South Carolina Press. At Google Books. Retrieved 11 August 2013.
  5. ^ Laird, Mark (1999) The Flowering of the Landscape Garden: English Pleasure Grounds, 1720-1800, p. 403. University of Pennsylvania Press At Google Books. Retrieved 11 August 2013.
  6. ^ "RHS Plant Selector - Salix 'Boydii'". Retrieved 2 June 2013. 
  7. ^ "Mourning Cloak". Study of Northern Virginia Ecology. Fairfax County Public Schools.
  8. ^ Albury/Wodonga Willow Management Working Group (December 1998). "Willows along watercourses: managing, removing and replacing". Department of Primary Industries, State Government of Victoria. 
  9. ^ Cremer, Kurt W. (2003). "Introduced willows can become invasive pests in Australia" (PDF). 
  10. ^ Salix spp. UFL/edu, Weeping Willow Fact Sheet ST-576, Edward F. Gilman and Dennis G. Watson, United States Forest Service
  11. ^ "Rooting Around: Tree Roots", Dave Hanson, Yard & Garden Line News Volume 5 Number 15, University of Minnesota Extension, October 1, 2003
  12. ^ Blackman, R. L.; Eastop, V. F. (1994). Aphids on the World's Trees. CABI. ISBN 9780851988771. 
  13. ^ David V. Alford (2012). Pests of Ornamental Trees, Shrubs and Flowers. p. 78. 
  14. ^ Kenaley, Shawn C. et al. (2010). "Leaf Rust" (PDF). 
  15. ^ James Breasted (English translation). "The Edwin Smith Papyrus". Retrieved 2007-06-09. 
  16. ^ "An aspirin a day keeps the doctor at bay: The world's first blockbuster drug is a hundred years old this week". Retrieved 2007-06-09. 
  17. ^ W. Hale White. "Materia Medica Pharmacy, Pharmacology and Therapeutics". Retrieved 2011-04-02. 
  18. ^ D. S. Vohra (1 June 2004). Bach Flower Remedies: A Comprehensive Study. B. Jain Publishers. p. 3. ISBN 978-81-7021-271-3. Retrieved 2 September 2013. 
  19. ^ "Flower remedies". Cancer Research UK. Retrieved September 2013. 
  20. ^ The palaeoenvironment of the Antrea Net Find The Department of Geography, University of Helsinki
  21. ^ Hageneder, Fred (2001). The Heritage of Trees. Edinburgh : Floris. ISBN 0-86315-359-3. p.172
  22. ^ Aylott, Matthew J.; Casella, E; Tubby, I; Street, NR; Smith, P; Taylor, G (2008). "Yield and spatial supply of bioenergy poplar and willow short-rotation coppice in the UK" (PDF). New Phytologist 178 (2): 358–370. doi:10.1111/j.1469-8137.2008.02396.x. PMID 18331429. Retrieved 2008-10-22. 
  23. ^ Mola-Yudego, Blas; Aronsson, Pär. (2008). "Yield models for commercial willow biomass plantations in Sweden" (PDF). Biomass and Bioenergy 32 (9): 829–837. doi:10.1016/j.biombioe.2008.01.002. Retrieved 2009-11-20. 
  24. ^ "Forestresearch.gov.uk". Forestresearch.gov.uk. Retrieved 2011-12-18. 
  25. ^ "ChurchYear.net". ChurchYear.net. Retrieved 2011-12-18. 
  26. ^ Doolittle, Justus (2002) [1876]. Social Life of the Chinese. Routledge. ISBN 978-0-7103-0753-8. 
  27. ^ Doré S.J., Henry; Kennelly, S.J. (Translator), M. (1914). Researches into Chinese Superstitions. Tusewei Press, Shanghai.  Vol I p. 2
  28. ^ "The Forest of Willows in Our Minds". Arirang TV. August 20, 2007. Retrieved September 10, 2007. 
  29. ^ "In Worship of Trees by George Knowles: Willow". 
  30. ^ "Mythology and Folklore of the Willow". 
  31. ^ "Under The Willow Tree". Hca.gilead.org.il. 2007-12-13. Retrieved 2011-12-18. 
  32. ^ "Green Willow". Spiritoftrees.org. Retrieved 2011-12-18. 
  33. ^ The Willow Wife[dead link]
  34. ^ "Wisdom of the Willow Tree". Tweedsblues.net. Retrieved 2011-12-18. 

Bibliography[edit]

  • Keeler, Harriet L. (1990). Our Native Trees and How to Identify Them. New York: Charles Scriber's Sons. Pages 393–395. ISBN 0-87338-838-0.
  • Newsholme, C. (1992). Willows: The Genus Salix. ISBN 0-88192-565-9
  • Warren-Wren, S.C. (1992). The Complete Book of Willows. ISBN 0-498-01262-X
  • Sviatlana Trybush, Šárka Jahodová, William Macalpine and Angela Karp. (2008). A genetic study of a Salix germplasm resource reveals new insights into relationships among subgenera, sections and species BioEnergy Research. 1(1):67 – 79.
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Salix bonplandiana

Salix bonplandiana (Bonpland willow), (Span.: ahuejote, sauce, ahujote, and huejote),[1] is a perennial species of willow tree native to southern and southwest Mexico and extending into central Guatemala;[2] in western Mexico it is a tree of the Sierra Madre Occidental cordillera, but also occurring in other small locales, for example Baja California Sur, northern Sonora, San Luis Potosi, etc. A core disjunct area occurs in central and southeast Arizona, in advantageous locales, especially associated with higher elevations and water.[3]

Distribution[edit]

In Mexico, the Bonpland willow is associated with the Pacific Coast, and in southern Mexico, the range extends into internal mountain areas of Pacific, central-southwest Guatemala. Across southern Mexico, it is a species of the Trans-Mexican Volcanic Belt,[4] and has its range extension on the Pacific, west from the west of the Volcanic Belt, and then north into the Sierra Madre Occidental cordillera. In the cordillera, the range ends in southwest Chihuahua, but is disjunct in a large area of central Arizona, the Mogollon RimWhite Mountains (Arizona) and extending into the Madrean Sky Islands southeastwards, including mountain-related isolated locales in the bootheel of extreme southwest New Mexico.[5]

Identical ranges of S. bonplandiana, S. taxifolia[edit]

Both S. bonplandiana, and S. taxifolia have identical ranges; both species range from the Trans-Mexican Volcanic Belt southward, on an imaginary centerline into central Guatemala. Each is on the Pacific Coast, but leave the coastal strip before the range extension into central Guatemala. Both have their disjunct ranges in Arizona, both being in the Madrean Sky Islands region. Both also range into Baja California Sur, but S. taxifolia only at the extreme south.[6] (S. bonplandiana also occurs in Arizona, in the Mogollon Rim–White Mountains regions.)

The other differences between the two are S. taxofolia occurs in minor locales of the Sierra Madre Oriental cordillera, and S. bonplandiana occurs in the Sierra Madre Occidental cordillera northwest to southwest Durango. S. taxifolia occurs west of the Occidental cordillera in the south Pacific Coast north to southern Sinaloa–southwest Durango. B. bonplandiana has a few locales on the Mexican Plateau, (with both occurring in central Chihuahua, west of Chihuahua, Chihuahua).[7] Other minor locale differences occur in central New Mexico, far west Texas, Coahuila, Nuevo Leon, and Tamaulipas.

Notes[edit]

  1. ^ City trees, (www.jornada.unam.mx); [1]
  2. ^ Little, Atlas of United States Trees, Volume 3, Minor Western Hardwoods, Map 166-SW, Map 166-N Salix bonplandiana.
  3. ^ Little, Map 166-SW, Map 166-N Salix bonplandiana.
  4. ^ Little, Map 166-SW, Map 166-N Salix bonplandiana.
  5. ^ Little, Map 166-SW, Map 166-N Salix bonplandiana.
  6. ^ Little, Maps 166-N & 183-N; Maps 166-SW & 183-SW, S. bonplandiana, S. taxifolia.
  7. ^ Little, Maps 166-N & 183-N; Maps 166-SW & 183-SW, S. bonplandiana, S. taxifolia.

References[edit]

  • Little. Atlas of United States Trees, Volume 3, Minor Western Hardwoods, Little, Elbert L, 1976, US Government Printing Office. Library of Congress No. 79-653298. Map 166-SW, Map 166-N Salix bonplandiana.
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Salix taxifolia

Salix taxifolia (yewleaf or yew-leaf willow) is a species of willow native to all of southern Mexico, also Pacific Coast regions, north to Sinaloa, and in the south Pacific Coast of Mexico into central Guatemala. Scattered populations are also reported from northern Mexico and from the US states of Texas, New Mexico, and Arizona. [3][4][5]

It is a large shrub or tree with narrow linear leaves similar to those of a yew (Taxus spp.), thus its common name. Its range is similar to that of the Bonpland willow, S. bonplandiana.

Distribution[edit]

The primary range of yewleaf willow is southern Mexico south of the Trans-Mexican Volcanic Belt and the Pacific coast region, then into Pacific coast–central Guatemala.[3]

Besides the core range area, (of the northern Sierra Madre Occidentals) in Arizona–New Mexico and northeast Sonora,[3] two larger disjunct regions occur in west Texas and central Chihuahua.[3] South of Chihuahua, Chihuahua it is found at the Conchos River, and west of the city, a large area at the lake region. It also occurs in scattered, isolated locales of Durango, Sinaloa, and in the northeast at Nuevo Leon and Tamaulipas: also extreme southern Baja California Sur, (west of Sinaloa-Durango across the Sea of Cortez).[3] Besides parts of the Sierra Madre Occidental, locales occur in the southern Sierra Madre Oriental cordillera, but also small isolated locales as far northeast as the northeast states.

Its range is similar to that of the Bonpland willow, Salix bonplandiana, except S. bonplandiana covers almost the entire north-south extent of the Sierra Madre Occidentals; yewleaf is more intermittent in that range.

Similar ranges of S. taxifolia, S. bonplandiana[edit]

Both S. taxifolia, and S. bonpandiana have identical ranges: both species range from the Trans-Mexican Volcanic Belt southward, on an imaginary centerline into central Guatemala. Each is on the Pacific Coast, but leave the coastal strip before the range into central Guatemala. Both have their disjunct ranges in Arizona, both being in the Madrean Sky Islands region. Both also range into Baja California Sur, but S. taxifolia only at the extreme south.[6] (S. bonplandiana also occurs in Arizona, in the Mogollon Rim–White Mountains regions.)

The other differences between the two: S. taxofolia occurs in minor locales of the Sierra Madre Oriental cordillera, and S. bonplandiana occurs in the Sierra Madre Occidental cordillera northwest to southwest Durango. S. taxifolia occurs west of the Occidental cordillera in the south Pacific Coast north to southern Sinaloa–southwest Durango. B. bonplandiana has a few locales on the Mexican Plateau, (with both occurring in central Chihuahua, west of Chihuahua, Chihuahua).[6] Other minor locale differences occur in central New Mexico, far west Texas, Coahuila, Nuevo Leon, and Tamaulipas.

References[edit]

  1. ^ Tropicos
  2. ^ The Plant List
  3. ^ a b c d e Elbert L. Little (1976), Atlas of United States Trees, Volume 3, Minor Western Hardwoods, US Government Printing Office. Library of Congress No. 79-653298, Map 183-SW, Map 183-N, Salix taxifolia.
  4. ^ Flora of North America, vol 7 p 54.
  5. ^ Laferrière, Joseph E. 1994. Vegetation and flora of the Mountain Pima village of Nabogame, Chihuahua, Mexico. Phytologia 77:102-140.
  6. ^ a b Little (1976), Maps 166-N & 183-N; Maps 166-SW & 183-SW, S. bonplandiana, S. taxifolia.
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