Comprehensive Description

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 Herb or subshrub to 50 cm, epipetric or terrestrial. Stems erect, drying brown to dark brown, green and maroon at the nodes when fresh, glabrous, brown peltate glandular on young stems and flowering nodes, cystoliths elliptic or absent, the internodes 18–200 × 1.5–2.0 mm, irregularly circular and grooved in cross-section, striate. Stipules 2.0–2.5 mm, deltate, drying red-brown. Leaves petiolate, distichous; petioles at each node unequal by ratio 1:1.8-8.5, 1.5–37 mm, glabrous; laminae at each node subequal or unequal by ratio 1: 1.9–5.4, laminae 12–130 × 5–26 mm, ovate, lanceolate, asymmetrically elliptic, oblanceolate or obovate, chartaceous; 3–nerved, the lateral nerves visible for >7/8 or more of the lamina length, secondary nerves 8-14 pairs, borne 60–75° to the midrib, weakly curved; upper surface drying dark brown, dark green when fresh, glabrous, cystoliths 0.250–0.375 mm, ‘V’ shaped, midrib raised; lower surface drying brown to dark brown, pale green when fresh, glabrous, eglandular; base symmetrical or asymmetrical, cuneate or obtuse; margin discretely serrulate or serrate, the basal 1/8–1/4 entire; apex symmetrical, attenuate to acuminate. Inflorescences ca 3 per stem, unisexual, staminate and pistillate inflorescences synchronous, born on separate stems; bracts 0.75–1.0 mm; bracteoles 0.50–0.675 mm. Staminate inflorescences 15–20 mm, bearing ca 60 flowers in a loose cyme; peduncle ≤1/4 inflorescence length, 0.5 mm in diameter, glabrous; pedicels 1.0–1.5 mm, glabrous. Staminate flowers 1.5 × 1.5 mm immediately prior to anthesis, pale green; tepals 4, 1.75 mm, valvate, broad elliptic or obovate, fused for their basal 1/2, glabrous, the subapical appendage ≤0.25 mm, ridge-like, glabrous; stamens 4. Pistillate inflorescences solitary, 2.0–3.5 mm, bearing 19-42 flowers in a compact cyme; peduncle 1/4 to 1/3 inflorescence length, 0.375 mm in diameter, glabrous, cystoliths absent; pedicels 0.375–0.50 mm, glabrous. Pistillate flowers 0.50–0.75 mm, tepals 3, unequal, adaxial tepal 0.5 mm, oblong to keel-shaped, the tepal appendage 0.375–0.50 mm, appearing inflated; the lateral tepals 0.375 mm, asymmetrically ovate; staminodes not visible. Infructescences 4.0–4.5 mm; achenes 0.675 × 0.375 mm, compressed, asymmetrically ovoid, the margin narrowly thickened.


Creative Commons Attribution 3.0 (CC BY 3.0)

© Alex K. Monro, Y. G. Wei, C. J. Chen

Source: PhytoKeys

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